A phylogenetic analysis of the Eucraniini (Coleoptera: Scarabaeidae: Scarabaeinae)

2002 ◽  
Vol 33 (3) ◽  
pp. 241-252 ◽  
Author(s):  
Federico C. Ocampo ◽  
Clarke H. Scholtz ◽  
T. Keith Philips

AbstractA phylogenetic analysis of the New World Eucraniini was conducted using cladistics. The tribe is monophyletic based on an analysis of 64 morphological characters. The flying Ennearabdus is the sister taxon to the three flightless genera, Anomiopsoides, Eucranium, and Glyphoderus, and Anomiopsoides and Glyphoderus share a common ancestor. The relationship of the eucraniines to other tribes of Scarabaeinae, dung preference, a comparison to the flightless southwest African genus Pachysoma, and the biogeography of the tribe are also discussed.

2004 ◽  
Vol 35 (1) ◽  
pp. 43-63 ◽  
Author(s):  
◽  
T. Keith Philips ◽  
Clarke Scholtz

AbstractA phylogenetic analysis of Phanaeini based on 137 morphological characters supports the hypothesis that the nine included genera, Coprophanaeus, Dendropaemon, Diabroctis, Homalotarsus, Megatharsis, Oxysternon, Phanaeus, Sulcophanaeus and Tetrameira, form a monophyletic clade. Monophyly is unaffected by the inclusion of Gromphas, Oruscatus, and Bolbites and these should also be considered phanaeines. The sister lineage is Ennearabdus (Eucraniini) and both evolved from ancestral Dichotomiini within South America. There is no support for a close relationship with the Onitini or any other remaining tribe. All phanaeine genera appear to be monophyletic except Sulcophanaeus, of which two species groups appear as sister taxa while the remaining three form an independent paraphyletic clade. Ancestral phanaeines were coprophagous with necrophagy evolving at least twice. Myrmecophily is also derived and most likely evolved only once in the common ancestor of Dendropaemon, Homalotarsus, Megatharsis and Tetramereia. Bare dung ball construction for larval development is also the most likely ancestral condition with a soil covering on the exterior ball surface and parental cooperation evolving in the more derived lineages.


Zootaxa ◽  
2021 ◽  
Vol 4926 (3) ◽  
pp. 401-416
Author(s):  
MOHAMMAD HUSSAIN FALAHZADAH ◽  
EBRAHIM SHOKOOHI ◽  
GHOLAM HOSSEIN MORAVEJ ◽  
PHATU WILLIAM MASHELA ◽  
ABDUL KHALID MADADI ◽  
...  

Several soil samples from different habitats in Badakhshan province of Afghanistan were collected to isolate and characterize bacteria feeding nematodes. The Galleria mellonella-baiting method was used for the isolation of the Afghan insect-associated nematodes. The nematodes were studied using morphological and morphometric data. The Oscheius specimen was characterized by a longer body (630–820 µm) and shorter pharynx (125–145 µm), whereas other morphological characters were not unusual. The Diploscapter specimen had an annulated cuticle, with lip region width 1.5 times shorter than the stoma, and had separated pharyngeal corpus from the isthmus and vulva located in the middle of the body. The molecular data were derived using three loci; 18S, 28S (D2/D3 segment), and ITS rRNA region, which were utilized to measure the genetic distance. The phylogenetic analysis was conducted to reconstruct the relationship tree. Both morphological and molecular approaches confirmed the identity of nematode isolates as Oscheius tipulae and Diploscapter coronatus. This is the first report of insect-associated nematodes from the soil of Afghanistan. Both species were capable of infecting and killing G. mellonella larvae in less than 96 h. 


2008 ◽  
Vol 38 (1-6) ◽  
pp. 129-175
Author(s):  
Christopher H. Dietrich ◽  
Dmitry A. Dmitriev

The 35 previously described species (including 4 junior synonyms) of the New World erythroneurine leafhopper genus Zyginama are reviewed and descriptions are provided for 43 new species, 2 from U.S.A., 26 from Mexico, 1 from Panama, and 14 from the Amazon region of South America. The following new synonyms are recognized: Z. aucta (McAtee) equals Erythroneura bilocularis Van Duzee, syn. n., and E. inclita Beamer, syn. n.; Z. nicholi (Beamer) equals E. canyonensis Beamer, syn. n., and E. ales Beamer, syn. n. Separate keys are provided for identification of males from North and South America, all known species are illustrated, and data on their distributions are summarized. Phylogenetic analysis of 40 morphological characters of the 71 species for which males are known yielded a reasonably well resolved estimate of relationships among the included species, but branch support was low overall.


2019 ◽  
pp. 54-78
Author(s):  
Yaroslav Tsytsiura

The article presents the results of a long-term study of the features of the phytocenology of agrophytocenoses of oil radish using different methods for assessing their condition and studying the tactics of the plant vitality strategy. A comprehensive assessment was made of the effect of changes in row spacing, seeding rates and fertilizer rates on the formation of various plant morphotypes, the variability of morphological characters and the general characteristics of plant viability. Three idiotypes of the plant oil radish were identified in a vertical study of agrophytocenosis, on the basis of which a detailed analysis of the variability of each group and a statistical assessment of the reliability of its existence were carried out. The features of the morphological integration of each tier are analyzed and its impact on the formation of the overall field productivity is evaluated. Based on the modular and vitality grouping, the effectiveness and feasibility of combining various options for plant density and fertilizer in the range of 30-90 kg of active substance per 1 ha were evaluated. Conclusions are drawn about the desired model of agrophytocenosis of oil radish based on the characteristics of its vital tactics and the goals of its growing. Grouping was carried out according to a variety of morphological traits of plants in the population and the possibilities of applying the basic patterns of phytocenology in their application to radish oilseed plants were evaluated. Through the use of regression analysis, the influence of climatic conditions on the formation of various morphological types of plants and the nature of the relationship of plants of oil radish in cenoses of various densities against the background of various fertilizer options has been evaluated. The main promising areas for further research on the peculiarities of creating highly productive and highly adaptable agrophytocenoses of oil radish have been outlined.


PeerJ ◽  
2018 ◽  
Vol 6 ◽  
pp. e5300
Author(s):  
Terry A. Gates ◽  
Khishigjav Tsogtbaatar ◽  
Lindsay E. Zanno ◽  
Tsogtbaatar Chinzorig ◽  
Mahito Watabe

We describe a new iguanodontian ornithopod,Choyrodon barsboldigen. et sp. nov. from the Albian-aged Khuren Dukh Formation of Mongolia based on several partial skeletons interpreted to represent a subadult growth stage based on osteohistological features. This new taxon is diagnosed by many autapomorphies of the maxilla, nasal, lacrimal, opisthotic, predentary, and surangular.Choyrodondisplays an unusual combination of traits, possessing an open antorbital fenestra (a primitive ornithopod trait) together with derived features such as a downturned dentary and enlarged narial fenestra. Histological imaging suggests that the type specimen ofChoyrodonwould have been a subadult at the time of death. Phylogenetic analysis of two different character matrices do not positChoyrodonto be the sister taxon or to be more primitive than the iguanodontianAltirhinus kurzanovi, which is found in the same formation. The only resolved relationship of this new taxon is that it was hypothesized to be a sister-taxon with the North American speciesEolambia caroljonesa. Though discovered in the same formation andChoyrodonbeing smaller-bodied thanAltirhinus, it does not appear that the former species is an ontogimorph of the latter. Differences in morphology and results of the phylogenetic analyses support their distinction although more specimens of both species will allow better refinement of their uniqueness.


1972 ◽  
Vol 104 (7) ◽  
pp. 1055-1071 ◽  
Author(s):  
Akira Mutuura

AbstractIn the phylogenetic classification of the Lepidoptera, the position of the genital opening and its modifications help to clarify the relationship of the Zeugloptera to the lepidopterous suborders. Three different systems of the suborder classification of the Lepidoptera, into Homoneura and Heteroneura (Tillyard, Imms, etc.), Monotrysia and Ditrysia (Borner), and Dacnonypha, Monotrysia, and Ditrysia (Hinton), are not supported by evidence obtained in the study of the female genitalia.The development of the female terminalia is closely associated with the mode of oviposition. Several types of female terminalia are found in the primitive Lepidoptera (Hepialidae, Neopseustidae, Tischeriidae, Lyonetiidae, Agathiphagidae, Tineidae, Nepticulidae, and Eriocraniidae). They are divided into four categories: hepialid type — the eggs are dropped randomly; typical lepidopterous type — the eggs are laid on the surface of food plants; eriocraniid type — the eggs are laid inside the food plants; tineid type — the eggs are laid in crevices of food. Each of the types is derivable from the types occurring in Neopseustidae, Tischeriidae, Agathiphagidae, and Eriocraniidae, respectively. These types are still far removed from the type of female terminalia in Micropterygidae of the Zeugloptera.The modes of egg-laying as well as the morphological characters of the female genitalia must be taken into consideration in suborder classification of the Lepidoptera.A genealogical tree based on the female terminalia suggests four main branches: hepialid branch — includes Neopseustidae and Prototheoridae; typical lepidopterous branch — includes all Ditrysia and Nepticulidae; tineid branch — includes only Tineidae; eriocraniid branch — includes Incurvariidae, Prodoxidae, Adelidae, and Heliozelidae.


1993 ◽  
Vol 125 (S168) ◽  
pp. 1-193 ◽  
Author(s):  
Valerie M. Behan-Pelletier

AbstractThe oribatid family Eremaeidae is represented in North America by two genera, Eremaeus and Eueremaeus, both widely distributed throughout the Palaearctic and Nearctic regions. In North America species in both genera are found in moist to arid habitats from New Mexico to the High Arctic. Reproduction is sexual, and both immatures and adults feed mainly on fungi.Revised diagnoses are presented for the Eremaeidae and genera Eremaeus and Eueremaeus. Eighteen species of Eremaeus, of which 14 are newly proposed, and 24 species of Eueremaeus, of which 15 are newly proposed, are recognized. Identification keys are provided for the world genera of Eremaeidae, and for adults of Eremaeus and Eueremaeus of North America. All but one North American species of these genera are described, and their geographical distributions mapped.North American Eremaeus species include E. appalachicus sp. no v., E. boreomontanus sp. nov., E. brevitarsus (Ewing), E. californiensis sp. nov., E. gracilis sp. nov., E. grandis Hammer, E. kananaskis sp. nov., E. kevani sp. nov., E. megistos sp. nov., E. monticolus sp. nov., E. nortoni sp. nov., E. occidentalis sp. nov., E. oresbios sp. nov., E. plumosus Woolley, E. porosus sp. nov., E. salish sp. nov., E. translamellatus Hammer, and E. walteri sp. nov. The immatures of four of these, E. kananaskis, E. occidentalis, E. oresbios, and E. translamellatus, are described.North American Eueremaeus include Eu. acostulatus sp. nov., Eu. aridulus sp. nov., Eu. columbianus (Berlese), Eu. foveolatus (Hammer), Eu. marshalli sp. nov., Eu. masinasin sp. nov., Eu. michaeli sp. nov., Eu. nahani sp. nov., Eu. nemoralis sp. nov., Eu. proximus (Berlese) comb, nov., Eu. woolleyi (Higgins) comb, nov., Eu. yukonensis sp. nov., and three informal species groups with the following included species in North America: (1) Eu. trionus group—Eu. trionus (Higgins) comb, nov., (2) Eu. stiktos group—Eu. carinatus sp. nov., Eu. higginsi sp. nov., Eu. stiktos (Higgins) comb, nov., Eu. tetrosus (Higgins) comb, nov., (3) Eu. chiatous group—Eu. alvordensis sp. nov., Eu. aysineep sp. nov., Eu. chiatous (Higgins) comb, nov., Eu. danos sp. nov., Eu. lindquisti sp. nov., Eu. magniporosus (Wallwork) comb, nov., and Eu. osoyoosensis sp. nov. The immatures of nine of these, Eu. masinasin, Eu. nahani, Eu. carinatus, Eu. higginsi, Eu. columbianus, Eu. proximus, Eu. woolleyi, Eu. stiktos, and Eu. tetrosus, are described. Kartoeremaeus reevesi Higgins and Eremaeus politus Banks are considered junior subjective synonyms of Eueremaeus columbianus (Berlese).A cladistic analysis of the genera comprising Eremaeidae: Eremaeus, Tricheremaeus, Eueremaeus (and included species groups), Proteremaeus, Carinabella, and Asperemaeus, indicates that Eremaeus is the sister taxon of Carinabella, and that Eueremaeus is the sister taxon of Proteremaeus. Tricheremaeus is the sister taxon of Eremaeus + Carinabella, and Asperemaeus is the sister taxon of Eueremaeus + Proteremaeus. The relationship of the Eremaeidae to the Megeremaeidae and Zetorchestidae is presented. Finally, I discuss the ecology and distribution of North American species of Eremaeidae.


2020 ◽  
Vol 50 (6) ◽  
Author(s):  
Elin A. Thomas ◽  
Ruoyu Liu ◽  
Diva Amon ◽  
Jon T. Copley ◽  
Adrian G. Glover ◽  
...  

AbstractChemosynthetic ecosystems have long been acknowledged as key areas of enrichment for deep-sea life, supporting hundreds of endemic species. Echinoderms are among the most common taxa inhabiting the periphery of chemosynthetic environments, and of these, chiridotid holothurians are often the most frequently observed. Yet, published records of chiridotids in these habitats are often noted only as supplemental information to larger ecological studies and several remain taxonomically unverified. This study therefore aimed to collate and review all known records attributed to Chiridota Eschscholtz, 1829, and to conduct the first phylogenetic analysis into the relationship of these chiridotid holothurians across global chemosynthetic habitats. We show that Chiridota heheva Pawson & Vance, 2004 is a globally widespread, cosmopolitan holothurian that occupies all three types of deep-sea chemosynthetic ecosystem—hydrothermal vents, cold seeps and organic falls—as an organic-enrichment opportunist. Furthermore, we hypothesise that C. heheva may be synonymous with another vent-endemic chiridotid, Chiridota hydrothermica Smirnov et al., 2000, owing to the strong morphological, ecological and biogeographical parallels between the two species, and predict that any chiridotid holothurians subsequently discovered at global reducing environments will belong to this novel species complex. This study highlights the importance of understudied, peripheral taxa, such as holothurians, to provide insights to biogeography, connectivity and speciation at insular deep-sea habitats.


Zootaxa ◽  
2011 ◽  
Vol 3047 (1) ◽  
pp. 1 ◽  
Author(s):  
MATTHEW S. WALLACE

Members of the Smiliini, the nominotypical tribe of the large New World subfamily Smiliinae, are predominately Nearctic in distribution. This tribe included 169 mostly tree-feeding species in 23 genera. A parsimony-based phylogenetic analysis of an original dataset comprising 89 traditional and newly discovered morphological characters for 69 species, including representatives of 22 of the 23 described genera of Smiliini and five other previously recognized tribes of the subfamily, resulted in a single most parsimonious tree with three major clades. The broad recent concept of Smiliini (including Telamonini as a junior synonym) was not recovered as monophyletic by the analysis. Instead, the analysis supported narrower definitions of both Telamonini, here reinstated from synonymy, and Smiliini. A key and diagnoses are given to define these tribes, along with discussions of their phylogeny, biogeography, and host plant associations. The genera Antianthe Fowler, Hemicardiacus Plummer, Smilirhexia McKamey, and Tropidarnis Fowler are placed as Smiliinae, incertae sedis. Based on the phylogeny, several genera from both tribes including Atymna Stål, Cyrtolobus Goding, Heliria Stål, and Telamona Fitch are not monophyletic. Diagnostic characters emphasizing the morphological differences between the Smiliini and Telamonini include the dorsal margin of the head, the shape of the pronotum, the size of the pronotal humeral angles, the presence or absence of pronotal longitudinal rugae, the size of forewing cells, variations in the fusion of veins R and M apically in both the foreand hind wing, and the shape of the apex of the female second valvulae. Mapping geographic distribution onto the phylogeny suggests that the common ancestor of the ingroup (all three clades) occurred in Central America and Mexico, with multiple dispersals to temperate North America. Many Smiliini and Telamonini feed on various species of oak (Quercus) and the close evolutionary association between these insects and their hosts is discussed.


1985 ◽  
Vol 104 (2) ◽  
pp. 429-433 ◽  
Author(s):  
H. A. Esechie

SummaryField and laboratory studies were carried out in 1981 and 1982 to investigate the relationship of stalk morphology and chemical composition to lodging resistance in maize in the rainforest zone of Nigeria.Morphological characters correlated with lodging were plant height, diameter and length of basal internode, thickness of rind and weight of 5 cm basal section. Lodging was negatively correlated with grain yield, and with the percentages of total nonstructural carbohydrate, protein and potassium in the stalks. Premature stalk senescence and rot were common in varieties susceptible to lodging. Lodging had no relationship with leaf area and number of days from sowing to flowering.


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