Competition among seedlings of perennial grasses, subterranean clover, white clover, and annual ryegrass in replacement series mixtures

2000 ◽  
Vol 51 (3) ◽  
pp. 377 ◽  
Author(s):  
G. M. Lodge

Seedlings of 3 perennial grasses, Danthonia linkii Kunthcv. Bunderra, D. richardsonii Cashmore cv. Taranna(wallaby grasses), and Phalaris aquatica L. cv. Sirosa,were each grown in replacement series mixtures with seedlings ofTrifolium repens L. (white clover),Trifolium subterraneum L. var. brachycalycinum (Katzn.et Morley) Zorahy & Heller cv. Clare (subterraneanclover), and Lolium rigidum L. (annual ryegrass). Plantswere sown 5 cm apart in boxes (45 by 29 by 20 cm) at a density of 307plants/m2. Maximum likelihood estimates were usedto derive parameters of a non-linear competition model using the dry matterweights of perennial grasses and competitors at 3 harvests, approximately 168,216, and 271 days after sowing. Intra-plant competition was examined inmonocultures of each species, grown at plant spacings of 2, 5, and 8 cm apartwith plants harvested at the above times.Competition occurred in all perennial grass–competitor mixtures, exceptin those of each perennial grass with white clover and thephalaris–subterranean clover mixture (Harvest 1) and those withD. richardsonii and phalaris grown with white clover(Harvest 2). For D. richardsonii (Harvests 1 and 2) andD. linkii (Harvest 1 only) grown with white clover andthe phalaris–subterranean clover (Harvest 1), the two species in themixture were not competing. In the phalaris–white clover mixture, eachspecies was equally competitive (Harvests 1 and 2). These differences incompetition and aggressiveness reflected differences in individual plantweights in monocultures where there was an effect (P < 0.05) of species ondry matter weight per box, but no significant effect of plant spacing.These data indicated that for successful establishment,D. richardsonii and D. linkiishould not be sown in swards with either subterranean clover or white clover,or where populations of annual ryegrass seedlings are likely to be high.Phalaris was more compatible with both white clover and subterranean clover,but aggressively competed with by annual ryegrass.

2000 ◽  
Vol 40 (6) ◽  
pp. 813 ◽  
Author(s):  
G. M. Lodge

Two experiments were conducted on plots sown in autumn 1992, at Tamworth in northern New South Wales. The first compared the establishment of 3 perennial grasses when sown as monocultures or with competitors in either broadcast-sown swards or alternate row-sown plots. Sowing rate and species of competitor were also examined as factors affecting perennial grass establishment. Perennial grasses were Austrodanthonia richardsonii (synonym Danthonia richardsonii) cv. Taranna, A. bipartita (synonym D. linkii) cv. Bunderra, and Phalaris aquatica cv. Sirosa. Competitors were Trifolium subterraneum var. brachycalycinum cv. Clare, T. repens cv. Haifa, and Lolium rigidum cv. Wimmera. In spring 1992, competitors were removed from 144 of the 288 plots to prevent them from seeding. A second experiment compared the longer-term (1993–96) dry matter yield and persistence of these perennial grasses under continuous grazing in plots were the competitor was present in year 1 (1992) or in all years (1992–96). In spring 1992, mean dry matter yield of perennial grass was higher (P<0.001) in row-sown plots then those sown by broadcasting. Mean dry matter yield of perennial grass was lowest at low sowing rate, but not significantly different at medium and high sowing rates (about 350 kg DM/ha). Compared with the monocultures, the presence of a competitor reduced mean perennial grass dry matter yields by 48, 69 and 85%, respectively for white clover, subterranean clover and annual ryegrass. Perennial grass plant numbers were highest (P<0.001) in the medium and high sowing rates of the monocultures and in white clover competitor plots and lowest (P<0.001) in all broadcast-sown plots, where annual ryegrass was the competitor. By spring 1996, white clover and annual ryegrass had declined to a low level in the pasture in all plots and the only major competitor was subterranean clover (1200 kg DM/ha, 40% plant frequency). Mean dry matter yields were highest (P<0.001) for Sirosa in 1993, but with dry conditions in 1994 and continuous grazing they were highest for Bunderra in all other years. The implications of these data for devising sowing strategies to maximise the establishment of perennial grasses and their long-term persistence in this environment are discussed.


1956 ◽  
Vol 7 (5) ◽  
pp. 367 ◽  
Author(s):  
WD Andrew ◽  
CA Neal-Smith

Over the period 1952-1954 there was no significant difference in the yield of herbage produced annually by the addition to a Phalaris tuberosa L.–Trifolium subterraneum L. pasture mixture, of any one of the following grasses: Agropyron obtusiusculum Lange., Bromus coloratus Steud., Bromus inermis Leyss., Dactylis glomerata L., or Festuca arundinacea Schreb. There were indications of a small change in seasonal production where certain grasses, notably D. glomerata, were included in the mixture. Over the 3-year period the proportion of the sown grass component, in the mixtures where either D. glomerata, B. inermis, or B. coloratus were included, increased by a greater amount than where the simple mixture of phalaris and subterranean clover was used. The addition of each grass also lessened the amount of P. tuberosa in the sown grass component of the yield. In the third year, despite the varying proportions of the phalaris and associated sown grass species, the mean population of the sown perennial grasses in each treatment did not differ significantly from the mean figure of 1.34 plants/sq. lk. The increased production of the sown grass yield component following the association of certain of the above species with P. tuberosa suggests that the latter does not fully exploit the environment. The principle of including another perennial grass when sowing phalaris and subterranean clover might have wide application as a means of combatting "phalaris staggers".


1972 ◽  
Vol 12 (57) ◽  
pp. 355 ◽  
Author(s):  
KFM Reed

August-born Corriedale lambs were weaned at ten weeks old and, for the following 12 months, were grazed on 16 plots. Each plot had been sown six years previously, to one of eight pasture mixtures which differed in their grass component, but not in their clover component. Trifolium perenne, Phalaris tuberosa, Dacylis glomerata, Trifolium repens, T . subterraneum, and T. fragiferum were sown. Hordem leporinum volunteered in most plots. Where perennial grass species were sown, Trifolium repens (white clover) contributed less than two per cent to the pasture present on the plots. For the two plots where perennial grass was not sown, white clover contributed up to 31 per cent. The total for all sown clover species present in the pasture varied from approximately 100 to 1000 kg ha-1, or approximately 3-30 per cent. Apart from the two plots where perennial qrass was not sown, the clover species present was mainly Trifolium subterraneum (subterranean clover). Average fleece weights varied from 3.1-4.9 kg on the different plots. The amount of sown clover present in the pasture accounted for 77 per cent of the between-plot variation in fleece weight and 66 per cent of the between-plot variation in annual liveweight gain. Measurements of pasture growth were made for four pasture mixtures. Total pasture growth showed differences of over 50 per cent due to pasture mixture, yet, when grazed at 20 weaners per hectare, differences in annual liveweight gain and fleece weight (10 months wool), due to pasture mixture, did not exceed 10 per cent. Annual liveweight gain and fleece weight were not correlated total pasture growth. However, pasture growth rate measured at monthly intervals, accounted for 43 per cent of the variation in mean daily liveweight gain during these individual months.


2019 ◽  
Vol 70 (9) ◽  
pp. 800
Author(s):  
John W. Piltz ◽  
Simon J. Flinn ◽  
Leslie A. Weston

Barley grass (Hordeum spp.) is a relatively short lived annual that provides high quality grazing early in the season, but its seed heads cause contamination of wool and carcasses, and may irritate the mouth, eyes and nose of sheep. Treatments were imposed on established subterranean clover (Trifolium subterraneum L.) annual pasture in the same plots for three consecutive years (2015 to 2017) to evaluate changes in barley grass content. Treatments included: grazing alone (G), herbicide followed by grazing (HG), or a forage conservation harvest in early October, late October or early November consistent with an early silage harvest (ES), late silage harvest (LS) or hay cut (H). Grazing plus herbicide markedly reduced (P &lt; 0.05) barley grass numbers compared with all other treatments, but increased (P &lt; 0.05) the growth of annual ryegrass (Lolium rigidum L.). ES reduced (P &lt; 0.05) barley grass and increased (P &lt; 0.05) subterranean clover compared with H, but broadleaf weed content benefitted by LS in contrast to either ES or H. Although herbicide application was the most effective method for barley grass control, forage harvest timing could be used to beneficially manipulate pasture composition.


1997 ◽  
Vol 37 (5) ◽  
pp. 531
Author(s):  
S. G. Clark ◽  
J. Li ◽  
A. M. Johnson ◽  
G. N. Ward ◽  
J. F. Chin

Summary. The seed banks of 6 subterranean clover (Trifolium subterraneum L.) cultivars were sampled at 3 abandoned cultivar evaluation experiments in south-western Victoria (Hamilton, Macarthur and Timboon). Two were var. yanninicum (Trikkala and Larisa) and 4 were var. subterraneum (Denmark, Goulburn, Karridale and Leura). Seed was sampled in late summer–early autumn 1993, between 6 and 10 years after the sites were established and between 3 and 7 years after the sites were abandoned. The collected seed was separated into black and white seed, and a sample of the black seed was grown in a glasshouse and identified as either belonging to the sown cultivar or belonging to another cultivar/genotype. All white seed was assumed to belong to the sown cultivar. The aim was to determine if these widely used cultivars were persisting under farm management conditions and competition from perennial grass, other subterranean clover and annual weeds. With some variation between sites all cultivars were found to be persisting satisfactorily. Over the 3 sites, white-seeded cultivars averaged 460 kg/ha and 6640 seeds/m2; black-seeded cultivars averaged 260 kg/ha and 5590 seeds/m2. Contamination with other subterranean clover cultivars/ecotypes was generally low, except at Macarthur where the white-seeded cultivars were heavily contaminated (Trikkala 39% pure; Larisa 57% pure) with black seeds highlighting the poor adaptation of var. yanninicum to light soil types. Of the black seeds, over the 3 sites, average contamination level was 13% and ranged from 41% (Goulburn at Macarthur) to 1% (Leura at Timboon). The study indicates that in south-western Victoria, all the recently released cultivars are likely to persist and be productive, and that the small-plot evaluation techniques used to identify potential new cultivars are likely to be adequate if replicated in space and if the clovers are sown in mixed swards with perennial grass.


2006 ◽  
Vol 46 (4) ◽  
pp. 439 ◽  
Author(s):  
W. McG. King ◽  
P. M. Dowling ◽  
D. L. Michalk ◽  
D. R. Kemp ◽  
G. D. Millar ◽  
...  

Temperate perennial grass-based pastures dominate the high rainfall zone of south-eastern Australia and support a major livestock production industry. This area has experienced a recent change in overall pasture condition, however, typified by a reduction in the abundance of perennial grasses and an increasingly prominent winter-annual grass weed component. Improving the condition and productivity of these pastures can be achieved by improved management but this requires better knowledge of the interactions between management options and pasture species composition and of the interaction between pasture vegetation and the complex effects of a heterogeneous landscape. This paper reports the results of an intensive survey of a 60-ha paddock that was designed to identify the species present, determine their patterns of distribution and examine the relationships between pasture vegetation and the environment. The survey of species present in late summer was supplemented by the identification of seedlings that later emerged from extracted soil cores and by soil physical and chemical analyses. Data were analysed using ordination and interpreted with GIS software so that topographic features could be considered. The most frequently identified taxa were Hypochaeris radicata, Austrodanthonia spp. and Bothriochloa spp. (in late summer) and Vulpia spp., Bromus molliformis and Trifolium subterraneum (winter-annual species). Austrodanthonia spp. were commonly found on the drier ridges and more acid soils with lower phosphate levels. These were also the areas dominated in spring by Vulpia spp. and were generally lower in plant species richness overall. The most species-rich areas occurred downslope where soil fertility was higher and less moisture stress was presumably experienced. The measured environmental factors explained a substantial proportion of the variation in the vegetation dataset, which underlined the importance of considering landscape effects in the management of typical tablelands pastures.


1960 ◽  
Vol 11 (3) ◽  
pp. 277 ◽  
Author(s):  
JN Black

Three strains of subterranean clover differing in leaf development were grown in pure swards and in all combinations in mixtures. The strain Yarloop has relatively few large leaves held on long petioles; Tallarook has many small leaves and short petioles; Bacchus Marsh is intermediate in all these respects. The swards were grown in boxes and were sampled on four occasions during the period of vegetative growth. Leaf area in each 2 cm layer of the sward was determined separately, and for both strains in the mixed swards, and measurements of the light intensity reaching each layer were obtained. Root weights were determined for all swards and separately for each component of the mixed swards. Examination of the dry weights showed that the growth of the strain having the lesser petiole elongation was suppressed. In the extreme instance Tallarook was so suppressed when grown with Yarloop that in the final inter-sampling period it grew completely in the dark, and lost about half its dry weight. Bacchus Marsh also suppressed Tallarook, but to a lesser extent, while Bacchus Marsh was itself suppressed in competition with Yarloop. The amount of light energy intercepted by the two components of the mixed swards was calculated from the leaf area and light profiles, and confirmed the importance of the spatial distribution of leaves in plant competition. It was concluded that, in the absence of defoliation, the success of a strain under competition was associated with its potential petiole elongation.


1991 ◽  
Vol 42 (5) ◽  
pp. 893 ◽  
Author(s):  
DC Edmeades ◽  
FPC Blamey ◽  
CJ Asher ◽  
DG Edwards

Ten temperate pasture legumes inoculated with appropriate rhizobia were grown for 31 days in flowing solution culture. Solution ionic strength was approximately 2700 8M and contained inorganic nitrogen (150 , 8M NO3-) only at the commencement of the experiment. Solution pH was maintained at 4.5, 5.0, 5.5 and 6.0. Also, five aluminium (Al) treatments were imposed, with nominal Al concentrations of 0, 3, 6, 12 and 24 8M (2.5, 7.1, 8.3, 11.2 and 24.7 8M Al measured) at pH 4.5. Solution pH <6 . 0 markedly reduced total dry mass (TDM) in all cultivars of white clover (Trifolium repens) cvv. 'Grasslands Pitau, Huia, G18 and Tahora' and red clover (Trifolium pratense) cvv. 'Grassland Turoa and Pawera', and to a lesser extent in the two subterranean clover (Trifolium subterraneum) cvv. 'Tallarook and Woogenellup'. In contrast, solution pH had no effect on the growth of Lotus corniculatus cv. Maitland, while Lotus pedunculatus cv. Maku grew best at pH 4.5. Lotus pedunculatus cv. Maku grew best in solution where the sum of the activities of the monomeric Al species {Alm} was maintained at 5.9 8M. The growth of all other species was decreased with Al in solution, a 50% reduction in TDM being associated with c. 6 8M {Alm] for white clover and subterranean clover, and c. 3 8M in red clover and Lotus corniculatus cv. Maitland.


2004 ◽  
Vol 44 (12) ◽  
pp. 1165 ◽  
Author(s):  
A. M. Bowman ◽  
W. Smith ◽  
M. B. Peoples ◽  
J. Brockwell

Total productivity and legume nitrogen fixation (N2 fixation) in dryland pastures were examined in a 2 year study (1999–2001) on 118 farms in central-western New South Wales. Pasture exclosure cages, placed at 217 on-farm sites, were harvested on 7 occasions and the foliage hand-sorted according to species in order to measure shoot dry matter (DM). The separated legume shoot material collected in spring 1999 (52 different legume samples) and 2000 (76 different legume samples) from a subset of representative pastures (41 cages on 28 different farms in 1999, 32 cages on 25 different farms in 2000) was also analysed for concentration of nitrogen (%N) and 15N natural abundance. These data were subsequently used to calculate the proportion of the legume shoot N derived from atmospheric N (%Ndfa), comparative measures of the relative efficiency of N2 fixation (kg N fixed/t DM accumulated) and the amounts of shoot N fixed (kg N/ha). The survey encompassed 8 common pasture types, and 5 others that were less common, ranging from native perennial grass pastures with little legume content to lucerne (Medicago sativa L.) pastures with and without companion clovers. Fifteen legume species were found in the pastures, some only occasionally. Lucerne and white clover (Trifolium repens L.) were the only perennials. Mean spring estimates of %Ndfa were similar in 1999 and 2000 for lucerne (72 and 81%, respectively), rose clover (T. hirtum All., 82 and 77%) and annual medics (Medicago spp., 89 and 86%). For the remaining 12 legume species, measures of %Ndfa ranged from 64 to 95% and averaged 83%. Shoot %N contents were greater for lucerne than for the other 14 legumes and this was reflected in the comparative measures of N2 fixation which ranged from 14.5 kg N/t DM for rose clover to 25.7 kg N/t DM for lucerne in 2000. The most productive pasture type comprised lucerne plus balansa clover [T. michelianum Savi var. balansae (Boiss.) Azn.], white clover or arrowleaf clover (T. vesiculosum Savi), but all pasture types that contained lucerne were highly productive. Spring was the most productive season and summer the least. Lucerne was overwhelmingly the most productive legume and was responsible for >83% of the fixed N in those pastures that contained both lucerne and other legumes. Lucerne productivity was approximately uniform throughout the year whereas, for other pastures, especially those based on rose clover or subterranean clover (T. subterraneum L.), there were sharp peaks in spring and little or no dry matter production over summer. The presence of lucerne in pastures significantly (P<0.05) reduced broadleaf weeds. It was concluded that, where there are requirements in central-western New South Wales agriculture for uniform forage production throughout the year and a high input of fixed N, lucerne is substantially superior to other species.


1992 ◽  
Vol 32 (8) ◽  
pp. 1061 ◽  
Author(s):  
AM Ridley ◽  
DR Coventry

Yield responses of 3 mixed grass-clover pastures [Phalaris aquatica L. cv. Sirosa (phalaris), Dactylis glomerata cv. Porto (cocksfoot), and annual grass based Trifolium subterraneum L. cv. Trikkala (subterranean clover) pastures] were measured over 5 soil pH treatments at 2 sites in Victoria. One site (Beechworth) was strongly acidic [pH(CaCl2) < 4.21 to a depth of 40 cm and contained high concentrations of soil aluminium (Al). At the other site (Lake Rowan), yield responses to lime application had been measured previously, but only in Al-sensitive wheat cultivars. At Beechworth, pasture yield responses to lime were not consistent but. when observed. occurred in autumn and winter in all 3 pasture types. Phalaris pastures showed yield increases more often than cocksfoot and annual pastures. Low magnesium and calcium concentrations may have limited dry matter production, although yields were reasonable on all treatments. Where lime was applied, growth responses may have been due to alleviation of Al toxicity. At Beechworth, pasture yield was increased where lime increased pH from 4.2 to 4.6 and decreased soil Al (measured in 10 mmol CaCl2/L) from 11 to <3 �g/g soil. Herbage manganese concentrations were not high in phalaris and subterranean clover, and cocksfoot manganese standards were not available. At Lake Rowan (pH 4.7, Al <1 �g/g), no growth responses to lime were seen in any pasture treatment, and annual grass based pastures sometimes had higher yields than phalaris and cocksfoot pastures. On strongly acidic soils such as at Beechworth, incorporation of lime prior to pasture establishment should be considered. Perennial grasses may reduce further soil degradation through acidification. Soil A1 concentrations are commonly lower in ley-cropping areas, and the inclusion of perennial grasses in ley pastures requires further evaluation.


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