Long-term persistence of subterranean clover Trifolium subterraneum L.) cultivars at three sites in south-western Victoria

1997 ◽  
Vol 37 (5) ◽  
pp. 531
Author(s):  
S. G. Clark ◽  
J. Li ◽  
A. M. Johnson ◽  
G. N. Ward ◽  
J. F. Chin
Keyword(s):  
Perennial Grass ◽  
Subterranean Clover ◽  
Late Summer ◽  
Trifolium Subterraneum ◽  
Contamination Level ◽  
Black And White ◽  
Small Plot ◽  
Black Seeds ◽  
Western Victoria ◽  
Annual Weeds

Summary. The seed banks of 6 subterranean clover (Trifolium subterraneum L.) cultivars were sampled at 3 abandoned cultivar evaluation experiments in south-western Victoria (Hamilton, Macarthur and Timboon). Two were var. yanninicum (Trikkala and Larisa) and 4 were var. subterraneum (Denmark, Goulburn, Karridale and Leura). Seed was sampled in late summer–early autumn 1993, between 6 and 10 years after the sites were established and between 3 and 7 years after the sites were abandoned. The collected seed was separated into black and white seed, and a sample of the black seed was grown in a glasshouse and identified as either belonging to the sown cultivar or belonging to another cultivar/genotype. All white seed was assumed to belong to the sown cultivar. The aim was to determine if these widely used cultivars were persisting under farm management conditions and competition from perennial grass, other subterranean clover and annual weeds. With some variation between sites all cultivars were found to be persisting satisfactorily. Over the 3 sites, white-seeded cultivars averaged 460 kg/ha and 6640 seeds/m2; black-seeded cultivars averaged 260 kg/ha and 5590 seeds/m2. Contamination with other subterranean clover cultivars/ecotypes was generally low, except at Macarthur where the white-seeded cultivars were heavily contaminated (Trikkala 39% pure; Larisa 57% pure) with black seeds highlighting the poor adaptation of var. yanninicum to light soil types. Of the black seeds, over the 3 sites, average contamination level was 13% and ranged from 41% (Goulburn at Macarthur) to 1% (Leura at Timboon). The study indicates that in south-western Victoria, all the recently released cultivars are likely to persist and be productive, and that the small-plot evaluation techniques used to identify potential new cultivars are likely to be adequate if replicated in space and if the clovers are sown in mixed swards with perennial grass.

Competition among seedlings of perennial grasses, subterranean clover, white clover, and annual ryegrass in replacement series mixtures

2000 ◽  
Vol 51 (3) ◽  
pp. 377 ◽  
Author(s):  
G. M. Lodge
Keyword(s):  
White Clover ◽  
Plant Competition ◽  
Perennial Grass ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Perennial Grasses ◽  
Maximum Likelihood Estimates ◽  
Annual Ryegrass ◽  
Replacement Series ◽  
Successful Establishment

Seedlings of 3 perennial grasses, Danthonia linkii Kunthcv. Bunderra, D. richardsonii Cashmore cv. Taranna(wallaby grasses), and Phalaris aquatica L. cv. Sirosa,were each grown in replacement series mixtures with seedlings ofTrifolium repens L. (white clover),Trifolium subterraneum L. var. brachycalycinum (Katzn.et Morley) Zorahy & Heller cv. Clare (subterraneanclover), and Lolium rigidum L. (annual ryegrass). Plantswere sown 5 cm apart in boxes (45 by 29 by 20 cm) at a density of 307plants/m2. Maximum likelihood estimates were usedto derive parameters of a non-linear competition model using the dry matterweights of perennial grasses and competitors at 3 harvests, approximately 168,216, and 271 days after sowing. Intra-plant competition was examined inmonocultures of each species, grown at plant spacings of 2, 5, and 8 cm apartwith plants harvested at the above times.Competition occurred in all perennial grass–competitor mixtures, exceptin those of each perennial grass with white clover and thephalaris–subterranean clover mixture (Harvest 1) and those withD. richardsonii and phalaris grown with white clover(Harvest 2). For D. richardsonii (Harvests 1 and 2) andD. linkii (Harvest 1 only) grown with white clover andthe phalaris–subterranean clover (Harvest 1), the two species in themixture were not competing. In the phalaris–white clover mixture, eachspecies was equally competitive (Harvests 1 and 2). These differences incompetition and aggressiveness reflected differences in individual plantweights in monocultures where there was an effect (P < 0.05) of species ondry matter weight per box, but no significant effect of plant spacing.These data indicated that for successful establishment,D. richardsonii and D. linkiishould not be sown in swards with either subterranean clover or white clover,or where populations of annual ryegrass seedlings are likely to be high.Phalaris was more compatible with both white clover and subterranean clover,but aggressively competed with by annual ryegrass.

Sustainable grazing systems for the Central Tablelands of New South Wales. 1. Agronomic implications of vegetation - environment associations within a naturalised temperate perennial grassland

2006 ◽  
Vol 46 (4) ◽  
pp. 439 ◽  
Author(s):  
W. McG. King ◽  
P. M. Dowling ◽  
D. L. Michalk ◽  
D. R. Kemp ◽  
G. D. Millar ◽  
...  
Keyword(s):  
Perennial Grass ◽  
Acid Soils ◽  
Late Summer ◽  
Trifolium Subterraneum ◽  
Moisture Stress ◽  
Recent Change ◽  
Perennial Grasses ◽  
Management Options ◽  
Eastern Australia ◽  
Winter Annual

Temperate perennial grass-based pastures dominate the high rainfall zone of south-eastern Australia and support a major livestock production industry. This area has experienced a recent change in overall pasture condition, however, typified by a reduction in the abundance of perennial grasses and an increasingly prominent winter-annual grass weed component. Improving the condition and productivity of these pastures can be achieved by improved management but this requires better knowledge of the interactions between management options and pasture species composition and of the interaction between pasture vegetation and the complex effects of a heterogeneous landscape. This paper reports the results of an intensive survey of a 60-ha paddock that was designed to identify the species present, determine their patterns of distribution and examine the relationships between pasture vegetation and the environment. The survey of species present in late summer was supplemented by the identification of seedlings that later emerged from extracted soil cores and by soil physical and chemical analyses. Data were analysed using ordination and interpreted with GIS software so that topographic features could be considered. The most frequently identified taxa were Hypochaeris radicata, Austrodanthonia spp. and Bothriochloa spp. (in late summer) and Vulpia spp., Bromus molliformis and Trifolium subterraneum (winter-annual species). Austrodanthonia spp. were commonly found on the drier ridges and more acid soils with lower phosphate levels. These were also the areas dominated in spring by Vulpia spp. and were generally lower in plant species richness overall. The most species-rich areas occurred downslope where soil fertility was higher and less moisture stress was presumably experienced. The measured environmental factors explained a substantial proportion of the variation in the vegetation dataset, which underlined the importance of considering landscape effects in the management of typical tablelands pastures.

Effect of perennial pasture species on surface soil moisture and early growth and survival of subterranean clover (Trifolium subterraneum L.) seedlings

1997 ◽  
Vol 48 (5) ◽  
pp. 683 ◽  
Author(s):  
B. S. Dear ◽  
P. S. Cocks
Keyword(s):  
Mineral Soil ◽  
Subterranean Clover ◽  
Late Summer ◽  
Soil Surface ◽  
Trifolium Subterraneum ◽  
Annual Grass ◽  
Growth And Survival ◽  
Eastern Australia ◽  
Standing Dead ◽  
Clover Seed

Subterranean clover seedling numbers and growth in swards containing 1 of 5 perennial pasture species [phalaris (Phalaris aquatica) cv. Sirolan, cocksfoot (Dactylis glomerata) cv. Currie, lucerne (Medicago sativa) cv. Aquarius, wallaby grass (Danthonia richardsonii) cv. Taranna, and lovegrass (Eragrostis curvula) cv. Consol] were compared with those in typical annual pastures and pure clover swards in the wheatbelt of eastern Australia. Presence of a perennial species or the volunteer annual grass (Eragrostis cilianensis) increased the rate of drying of the soil surface (0–5 cm) after late February and May rain, compared with subterranean clover swards. Perennials differed in the rate they dried the soil surface, with the more summer-active lucerne and consul lovegrass drying the profile more rapidly than phalaris. The amount of water in the surface 5 cm, 6 days after the rainfall event on 27–28 February, was strongly negatively correlated (r = –0·75, P < 0·01) with the amount of green perennial biomass, but not related to standing dead material or surface residues. Where perennials were present, a smaller proportion (2–4%) of the clover seed pool produced seedlings in response to late summer rain, compared with pure clover swards (18%). A higher proportion of the seed pool produced seedlings (19–36%) following rain in late autumn but there was no difference between species. The more summer-active perennials (cocksfoot, danthonia, and lucerne) markedly depressed the survival of emerged clover seedlings following both germinations. Of the seedlings that emerged in early March, the proportion remaining by 29 March was 57% in phalaris, 21% in lucerne, 13% in danthonia, and 1% in cocksfoot, compared with a 78% increase in seedlings in pure subterranean clover swards. By 15 May, all perennials had <2 clover seedlings/m2 surviving, compared with 37 in the annual pasture and 964 plants/m2 in pure subterranean clover. Following the May germination, the highest proportion of emerged seedlings surviving until 29 May was in the phalaris swards (40%) and least in the cocksfoot and danthonia swards (2–4%). Presence of a perennial or annual grass decreased (P < 0·05) relative water content of clover seedlings on 15 March from 74% in pure clover swards, to 48% in annual pasture, 34% in phalaris, and 29% in lucerne swards. Clover seedlings growing in pure subterranean swards on 15 March (17 days after germinating rain) were 4 times larger than those in lucerne and twice as large as those in either phalaris or annual pasture. Seed size did not differ between treatments, but available mineral soil nitrogen was significantly higher (P < 0·001) in pure subterranean clover swards (32 mg N/g) compared with perennials (3–13 mg N/g). Strategies such as heavy grazing in late summer to reduce green biomass of the perennials or sowing the perennials at lower densities may reduce the adverse effects that perennials have on subterranean clover seedlings in these drier environments.

Phosphorus, sulphur, and molybdenum deficiencies in soils from eastern Australia in relation to nutrient supply and some characteristics of soil and climate.

1955 ◽  
Vol 6 (5) ◽  
pp. 673 ◽  
Author(s):  
KD McLachlan
Keyword(s):  
Phosphorus Deficiency ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Parent Material ◽  
The Other ◽  
Podzolic Soils ◽  
Sulphur Deficiency ◽  
Eastern Australia ◽  
Yield Responses ◽  
Western Victoria

Subterranean clover (Trifolium subterraneum L.) was grown on 32 virgin soils in pot cultures, and the yield responses to phosphorus, sulphur, and molybdenum were determined for each soil. The soils were collected from a wide area in eastern Australia, extending from south-western Victoria to southern Queensland. Sulphur deficiency occurred almost as frequently as phosphorus deficiency. In fact, 75 per cent. of the soils were deficient in both phosphorus and sulphur. Thirty-one per cent. were deficient in all three elements. The effect of the interaction between the elements on the occurrence and intensity of the deficiencies is shown. Molybdenum responses were obtained only after the other deficiencies had been corrected. A deficiency in one of the elements is no indication of deficiency or sufficiency of either of the other two. There was no correlation between the occurrence or intensity of the deficiencies and the geological origin of the soil parent material, the climate of the regions from which the soils were collected, or such soil characters as colour, organic matter, and texture. Responses to phosphorus were less on the black earths than on the red or yellow podzolic soils; those on the red earths were intermediate. The intensity of sulphur deficiency increased, and the intensity of molybdenum deficiency decreased, with increasing soil pH.

Tactical versus continuous stocking for persistence of perennial ryegrass (Lolium perenne L.) in pastures grazed by sheep in south-western Victoria

1999 ◽  
Vol 39 (3) ◽  
pp. 265 ◽  
Author(s):  
R. A. Waller ◽  
P. E. Quigley ◽  
G. R. Saul ◽  
G. A. Kearney ◽  
P. W. G. Sale
Keyword(s):  
Lolium Perenne ◽  
Perennial Ryegrass ◽  
Seedling Recruitment ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Fold Increase ◽  
Early Summer ◽  
Lolium Perenne L ◽  
Western Victoria ◽  
Tiller Density

The survival of perennial ryegrass (Lolium perenne L.) plants was studied in sheep pastures in south-western Victoria during the dry summer of 1996–97. Recruitment of perennial ryegrass seedlings into the pasture sward was also monitored in the autumn–winter periods in 1997 and 1998. The objective was to investigate whether a tactical stocking strategy, consisting of variable length summer, autumn and winter rotations and continuous stocking in spring, might increase perennial ryegrass tiller survival and seedling recruitment in the autumn, compared with continuous stocking all year. The grazing strategies were compared on 2 contrasting pastures: an upgraded pasture [sown with newer cultivars of perennial ryegrass and subterranean clover (Trifolium subterraneum L.) with 26 kg phosphorus/ha.year as applied fertiliser] and a naturalised perennial ryegrass pasture receiving 6 kg P/ha.year. Paddocks were grazed by Border Leicester × Merino ewes, mated to a terminal sire so as to lamb in September. Perennial ryegrass tiller density was higher on the upgraded pasture with a mean density of 7750 tillers/m2 in early summer which declined to zero live tillers by mid summer. Live tillers began to reappear before the opening rains and then increased after the rain. Mean tiller density in the upgraded pasture declined over the 2 summers, with only 2050 tillers/m2 being present 2 months after the opening rains in 1998. There were no effects (P>0.05) of pasture type or grazing strategy on the number of tagged tillers that survived the summer period. Only 12% of the vegetative tillers, randomly tagged in December 1996, survived to May 1997. More than half of the tillers (56%) that produced a seedhead produced daughter tillers which survived the dry summer–autumn period. A significant (P<0.05) interaction between grazing strategy and pasture type occurred with the number of perennial ryegrass seedlings that had established 4 weeks after the opening rains in 1997. There was a 5–11-fold increase in seedling numbers which regenerated in the tactically stocked, upgraded pasture compared with the other treatments. Seedling recruitment was considerably lower in the autumn of 1998, due presumably to an overall decline in perennial ryegrass density relative to annual grasses in 1997. A second experiment investigated the effect of excluding sheep from grazing at anthesis until seedhead maturation or until the opening rains, together with a mechanical seed dislodgment treatment at seed maturity. All exclusion treatments increased seedling recruitment 4–7-fold, compared with continuous stocking. The results suggest a possible mechanism by which perennial ryegrass density can be increased without expensive reseeding.

The effect of associate perennial grasses on a simple pasture mixture of phalaris and subterranean clover

1956 ◽  
Vol 7 (5) ◽  
pp. 367 ◽  
Author(s):  
WD Andrew ◽  
CA Neal-Smith
Keyword(s):  
Festuca Arundinacea ◽  
Perennial Grass ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Yield Component ◽  
Perennial Grasses ◽  
Grass Species ◽  
Significant Difference ◽  
The Mean ◽  
Small Change

Over the period 1952-1954 there was no significant difference in the yield of herbage produced annually by the addition to a Phalaris tuberosa L.–Trifolium subterraneum L. pasture mixture, of any one of the following grasses: Agropyron obtusiusculum Lange., Bromus coloratus Steud., Bromus inermis Leyss., Dactylis glomerata L., or Festuca arundinacea Schreb. There were indications of a small change in seasonal production where certain grasses, notably D. glomerata, were included in the mixture. Over the 3-year period the proportion of the sown grass component, in the mixtures where either D. glomerata, B. inermis, or B. coloratus were included, increased by a greater amount than where the simple mixture of phalaris and subterranean clover was used. The addition of each grass also lessened the amount of P. tuberosa in the sown grass component of the yield. In the third year, despite the varying proportions of the phalaris and associated sown grass species, the mean population of the sown perennial grasses in each treatment did not differ significantly from the mean figure of 1.34 plants/sq. lk. The increased production of the sown grass yield component following the association of certain of the above species with P. tuberosa suggests that the latter does not fully exploit the environment. The principle of including another perennial grass when sowing phalaris and subterranean clover might have wide application as a means of combatting "phalaris staggers".

Tactical versus continuous stocking in perennial ryegrass-subterranean clover pastures grazed by sheep in south-western Victoria. 2. Ryegrass persistence and botanical composition

2001 ◽  
Vol 41 (8) ◽  
pp. 1109 ◽  
Author(s):  
R. A. Waller ◽  
P. W. G. Sale ◽  
G. R. Saul ◽  
G. A. Kearney
Keyword(s):  
Perennial Ryegrass ◽  
Dry Matter ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Summer Rainfall ◽  
Grazing Pressure ◽  
Botanical Composition ◽  
Average Mass ◽  
Gravelly Soil ◽  
Western Victoria

A 4-year field experiment was carried out in south-western Victoria to determine whether tactical stocking might improve perennial ryegrass (Lolium perenne L.) persistence and prime lamb production, compared with the more common practice of continuous stocking. Tactical stocking consisted of variable-length summer, autumn and winter rotations and continuous stocking in spring. The 2 grazing strategies were compared on 2 contrasting pastures: an upgraded pasture, sown with newer cultivars of perennial ryegrass and subterranean clover (Trifolium subterraneum L.) with 26 kg phosphorus/ha.year, and a more typical naturalised perennial ryegrass pasture receiving 6 kg phosphorus/ha.year. Paddocks were grazed by Border Leicester x Merino ewes, which were mated to a terminal sire to lamb in September. The effects of the grazing systems and pasture treatments on ryegrass persistence and pasture botanical composition are presented in this paper. The perennial ryegrass component of the sward decreased from an average mass of 1000 kg dry matter/ha on the upgraded pasture and 400 kg dry matter/ha on the typical pasture in spring 1996 to 100 kg dry matter/ha and <50 kg/ha for the 2 pasture types, respectively, in spring 1998. This occurred irrespective of the grazing system imposed. The decline of ryegrass was attributed to several factors including the very dry 3-year period, summer rainfall events that caused ryegrass buds to break dormancy then die when dry conditions resumed, the gravelly soil type where the experiment was located, and the high grazing pressure used. A number of species became more prevalent in the pastures as the ryegrass disappeared. Silver grass (Vulpia spp.) increased 4–5-fold on all treatments. By the fourth year, barley grass (Hordeum leporinum) and Bromus spp. had increased significantly with tactical stocking compared with continuous stocking. Capeweed (Arctotheca calendula L.) and Erodium spp. increased on both the continuously stocked and upgraded pastures. The subterranean clover component was maintained across all treatments over the 4 years, indicating that tactical stocking did not adversely affect clover persistence. Since ryegrass declined irrespective of treatment, the tactical stocking system used in this study cannot be recommended for improving ryegrass persistence.

Tactical versus continuous stocking in perennial ryegrass-subterranean clover pastures grazed by sheep in south-western Victoria. 1. Stocking rates and herbage production

2001 ◽  
Vol 41 (8) ◽  
pp. 1099 ◽  
Author(s):  
R. A. Waller ◽  
P. W. G. Sale ◽  
G. R. Saul ◽  
G. A. Kearney
Keyword(s):  
Perennial Ryegrass ◽  
Dry Matter ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Stocking Rate ◽  
Soil Fertility Status ◽  
Western Victoria ◽  
Terminal Sire ◽  
Autumn And Winter ◽  
Stocking Rates

A 4-year field experiment was carried out in south-western Victoria to determine whether tactical stocking might improve perennial ryegrass (Lolium perenne L.) persistence and prime lamb production, compared with the more common practice of year-around continuous stocking. Tactical stocking consisted of variable length summer, autumn and winter rotations and continuous stocking in spring. The 2 grazing strategies were compared on 2 contrasting pastures: an upgraded pasture, sown with newer cultivars of perennial ryegrass and subterranean clover (Trifolium subterraneum L.) with 26 kg phosphorus/ha.year, and a more typical naturalised perennial ryegrass pasture receiving 6 kg phosphorus/ha.year. Paddocks were grazed by Border Leicester x Merino ewes, which were mated to a terminal sire to lamb in September. The effects of the grazing systems and pasture treatments on herbage production and stocking rate are presented in this paper. Herbage production was similar between the treatments, but tactical stocking significantly increased herbage mass during the growing season (P<0.05) compared with continuous stocking. In spring each year, the herbage mass generally exceeded 3000 kg dry matter/ha in tactically stocked paddocks and averaged 500–900 kg dry matter/ha higher than the mass on continuously stocked paddocks. This enabled the year-round stocking rate to be increased by an average of 9% over the 4 years of the experiment. We considered that the stocking rates could not be further increased, despite the higher herbage mass in spring, as stock reduced the dry herbage to a low residual mass by the opening rains in autumn. In contrast, stocking rates averaged 51% higher on the upgraded pasture compared with the typical pasture over the 4 years of the experiment. This indicates that pasture improvement and soil fertility status have a much greater impact on productivity than changes to grazing method. However, tactical stocking was able to increase the sustainability of prime lamb production on upgraded pastures in a dry summer climate, by maintaining herbage cover on the paddocks over the summer–autumn period.

Effect of grazing strategy, ryegrass overdrilling and herbicide application on vulpia content, tiller density and seed production in perennial pastures

2008 ◽  
Vol 48 (5) ◽  
pp. 632 ◽  
Author(s):  
K. N. Tozer ◽  
D. F. Chapman ◽  
P. E. Quigley ◽  
P. M. Dowling ◽  
R. D. Cousens ◽  
...  
Keyword(s):  
Soil Seed Bank ◽  
Subterranean Clover ◽  
Trifolium Subterraneum ◽  
Integrated Approach ◽  
Annual Rainfall ◽  
Herbicide Application ◽  
Effective Strategies ◽  
Annual Grasses ◽  
Western Victoria ◽  
Tiller Density

Vulpia species C.C. Gmel. are annual grass weeds that can reduce the productivity of perennial pastures throughout southern Australia. To develop more effective strategies to manage vulpia, a 3-year experiment was established in western Victoria (average annual rainfall: 625 mm) comparing different methods currently used to control this weed. Overdrilling perennial ryegrass (Lolium perenne L.) seed and simazine application treatments were applied to phalaris (Phalaris aquatica L.) pastures that were set-stocked or rotationally grazed (either as a four-paddock or strategic rotation) with Merino ewes. The content of vulpia, subterranean clover (Trifolium subterraneum L.) and other annual grasses as a proportion of total dry matter increased, and the proportion of phalaris decreased in most grazing treatments throughout the experiment. The mean vulpia content was lowest and the phalaris content was highest in the four-paddock rotation, whereas vulpia content was greatest and phalaris content was lowest under set-stocking. Simazine application in June with or without ryegrass overdrilling reduced the number of vulpia tillers/m2 in 2000 and 2001 and vulpia panicle production in 2000, although vulpia populations increased to pretreatment levels in herbicide-treated swards by 2002. The number of vulpia seeds in the soil seed bank was not affected by any of the treatments. The most effective treatment was a combination of ryegrass overdrilling and herbicide application in the four-paddock, rotationally grazed pastures. This experiment highlights the need for an integrated approach to manage vulpia since relying on herbicide application alone is ineffective. This is particularly the case when competitive pasture species are unable to adequately utilise available resources and prevent a recovery in vulpia populations.

Long term patterns of seed softening and seedling establishment from single seed crops of subterranean clover

1984 ◽  
Vol 24 (125) ◽  
pp. 200 ◽  
Author(s):  
GB Taylor ◽  
RC Rossiter ◽  
MJ Palmer
Keyword(s):  
Seedling Establishment ◽  
Subterranean Clover ◽  
Late Summer ◽  
Trifolium Subterraneum ◽  
The Other ◽  
Hard Seed ◽  
Single Seed ◽  
One Year ◽  
Seed Crops

Single seed crops of eight varieties of subterranean clover (Trifolium subterraneum) were grown in swards in four successive years in a medium rainfall environment at Bakers Hill, Western Australia. Measurements were made of seed production, the progress of seed softening within the first two summers, and residual hard seed and seedling numbers after the break of season for up to six successive regeneration years. Patterns of seed softening were also measured in a laboratory oven with a diurnal temperature fluctuation of 60/15�C. Approximately half the seeds of all varieties softened over the first summer and generally well over half the residual seeds softened during the second summer. Up to 18% of seeds of the hardest seeded variety, Northam, and 11% of Geraldton seeds persisted to the end of the third summer as hard seeds. Seed softening patterns showed a significant variety x regeneration year interaction due mainly to a slower rate of softening of Northam seeds relative to the other varieties after the first summer. In six of the varieties, seeds generally softened more slowly in the laboratory than in the field but in the case of the other two varieties this trend was reversed. Nevertheless, ranking of varieties for rate of seed softening was similar in field and laboratory. Seedling establishment ranged from 28 to 100% of soft seeds, depending on the incidence of late summer and autumn rains. It appeared that all varieties in the study would regenerate satisfactorily at densities of more than 15 plants/dm2 after at least one year in which little or no seed is set. Slower rates of softening would probably be desirable in drier regions or in rotation systems involving cropping.

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