scholarly journals Performance of steer progeny of sires differing in genetic potential for fatness and meat yield following postweaning growth at different rates. 2. Carcass traits

2009 ◽  
Vol 49 (6) ◽  
pp. 525 ◽  
Author(s):  
W. A. McKiernan ◽  
J. F. Wilkins ◽  
J. Irwin ◽  
B. Orchard ◽  
S. A. Barwick
Keyword(s):  
Production Systems ◽  
Carcass Traits ◽  
Quality Traits ◽  
Muscle Area ◽  
Eye Muscle ◽  
Meat Yield ◽  
Breeding Values ◽  
Genetic Potential ◽  
Estimated Breeding Values ◽  
Selection Of

The steer progeny of sires genetically diverse for fatness and meat yield were grown at different rates from weaning to feedlot entry and effects on growth, carcass and meat-quality traits were examined. The present paper, the second of a series, reports the effects of genetic and growth treatments on carcass traits. A total of 43 sires, within three ‘carcass class’ categories, defined as high potential for meat yield, marbling or both traits, was used. Where available, estimated breeding values for the carcass traits of retail beef yield (RBY%) and intramuscular fat (IMF%) were used in selection of the sires, which were drawn from Angus, Charolais, Limousin, Black Wagyu and Red Wagyu breeds, to provide a range of carcass sire types across the three carcass classes. Steer progeny of Hereford dams were grown at either conventional (slow: ~0.5 kg/day) or accelerated (fast: ~0.7 kg/day) rates from weaning to feedlot entry weight, with group means of ~400 kg. Accelerated and conventionally grown groups from successive calvings were managed to enter the feedlot at similar mean feedlot entry weights at the same time for the 100-day finish under identical conditions. Faster-backgrounded groups had greater fat levels in the carcass than did slower-backgrounded groups. Dressing percentages and fat colour were unaffected by growth treatment, whereas differences in ossification score and meat colour were explained by age at slaughter. There were significant effects of sire type for virtually all carcass traits measured in the progeny. Differences in hot standard carcass weight showed a clear advantage to European types, with variable outcomes for the Angus and Wagyu progeny. Sire selection by estimated breeding values (within the Angus breed) for yield and/or fat traits resulted in expected differences in the progeny for those traits. There were large differences in both meat yield and fatness among the types of greatest divergence in genetic potential for those traits, with the Black Wagyu and the Angus IMF clearly superior for IMF%, and the European types for RBY%. The Angus IMF progeny performed as well as that of the Black Wagyu for all fatness traits. Differences in RBY% among types were generally reflected by similar differences in eye muscle area. Results here provide guidelines for selecting sire types to target carcass traits for specific markets. The absence of interactions between growth and genetic treatments ensures that consistent responses can be expected across varying management and production systems.

scholarly journals Erratum to: Performance of steer progeny of sires differing in genetic potential for fatness and meat yield following postweaning growth at different rates. 2. Carcass traits

2009 ◽  
Vol 49 (11) ◽  
pp. 1052
Author(s):  
W. A. McKiernan ◽  
J. F. Wilkins ◽  
J. Irwin ◽  
B. Orchard ◽  
S. A. Barwick
Keyword(s):  
Production Systems ◽  
Carcass Traits ◽  
Quality Traits ◽  
Muscle Area ◽  
Eye Muscle ◽  
Meat Yield ◽  
Breeding Values ◽  
Genetic Potential ◽  
Estimated Breeding Values ◽  
Selection Of

The steer progeny of sires genetically diverse for fatness and meat yield were grown at different rates from weaning to feedlot entry and effects on growth, carcass and meat-quality traits were examined. The present paper, the second of a series, reports the effects of genetic and growth treatments on carcass traits. A total of 43 sires, within three 'carcass class' categories, defined as high potential for meat yield, marbling or both traits, was used. Where available, estimated breeding values for the carcass traits of retail beef yield (RBY%) and intramuscular fat (IMF%) were used in selection of the sires, which were drawn from Angus, Charolais, Limousin, Black Wagyu and Red Wagyu breeds, to provide a range of carcass sire types across the three carcass classes. Steer progeny of Hereford dams were grown at either conventional (slow: ~0.5 kg/day) or accelerated (fast: ~0.7 kg/day) rates from weaning to feedlot entry weight, with group means of ~400 kg. Accelerated and conventionally grown groups from successive calvings were managed to enter the feedlot at similar mean feedlot entry weights at the same time for the 100-day finish under identical conditions. Faster-backgrounded groups had greater fat levels in the carcass than did slower-backgrounded groups. Dressing percentages and fat colour were unaffected by growth treatment, whereas differences in ossification score and meat colour were explained by age at slaughter. There were significant effects of sire type for virtually all carcass traits measured in the progeny. Differences in hot standard carcass weight showed a clear advantage to European types, with variable outcomes for the Angus and Wagyu progeny. Sire selection by estimated breeding values (within the Angus breed) for yield and/or fat traits resulted in expected differences in the progeny for those traits. There were large differences in both meat yield and fatness among the types of greatest divergence in genetic potential for those traits, with the Black Wagyu and the Angus IMF clearly superior for IMF%, and the European types for RBY%. The Angus IMF progeny performed as well as that of the Black Wagyu for all fatness traits. Differences in RBY% among types were generally reflected by similar differences in eye muscle area. Results here provide guidelines for selecting sire types to target carcass traits for specific markets. The absence of interactions between growth and genetic treatments ensures that consistent responses can be expected across varying management and production systems.

scholarly journals Genetic Analysis of Major Carcass Traits of Korean Hanwoo Males Raised for Thirty Months

Animals ◽  
2021 ◽  
Vol 11 (6) ◽  
pp. 1792
Author(s):  
Mahboob Alam ◽  
Soo Hyun Lee ◽  
Do Hyun Lee ◽  
Chungil Cho ◽  
Mi Na Park
Keyword(s):  
Genetic Parameters ◽  
Carcass Traits ◽  
Production Cycle ◽  
Marbling Score ◽  
Multiple Trait ◽  
Muscle Area ◽  
Eye Muscle ◽  
Future Studies ◽  
Genetic Potential ◽  
First Time

Understanding animals’ genetic potential for carcass traits is the key to genetic improvements in any beef cattle. In this study, we investigated the genetic merits of carcass traits using Hanwoo males raised in a 30-month production system (PROD30). We achieved this using a dataset comprising 6092 Hanwoo males born between 2005 and 2017 and measures of four carcass traits (carcass weight, CWT; eye muscle area, EMA; backfat thickness, BFT; and marbling score, MS). Genetic parameters were estimated using a multiple-trait animal model through the AIREMLF90 program. According to the multiple-trait model, the h2 of CWT, EMA, BFT, and MS were 0.35 ± 0.04, 0.43 ± 0.05, 0.48 ± 0.05, and 0.56 ± 0.05, respectively. The strongest genetic correlation (rg) was obtained between CWT and EMA (0.49 ± 0.07), whereas it was negligible between CWT and BFT. EMA and MS were also moderately correlated, whereas there was a relatively low negative correlation between EMA and BFT (−0.26 ± 0.08). Our study revealed a consistent indirect genetic improvement in animals from 2005 onwards. Although Hanwoo improvement has mainly focused on males under a 24-month production cycle, we observed PROD30 males to have somewhat similar genetic potential. Our results provide useful insights into the genetic merits of PROD30 males for the first time, which may facilitate future studies on them and their integration into the Hanwoo National Evaluation for genomic selection.

Genotype and age effects on sheep meat production. 2. Carcass quality traits

2007 ◽  
Vol 47 (10) ◽  
pp. 1147 ◽  
Author(s):  
E. N. Ponnampalam ◽  
D. L. Hopkins ◽  
K. L. Butler ◽  
F. R. Dunshea ◽  
R. D. Warner
Keyword(s):  
Muscle Development ◽  
Carcass Weight ◽  
Meat Production ◽  
Carcass Quality ◽  
Quality Traits ◽  
Muscle Area ◽  
Muscle Weight ◽  
Eye Muscle ◽  
Younger Age ◽  
Estimated Breeding Values

The present study examines carcass quality traits in different genotypes of Australian sheep slaughtered at 4, 8, 14 and 22 months of age. Poll Dorsetgrowth × Border Leicester Merino (PDg × BLM), Poll Dorsetgrowth × Merino (PDg × M), Poll Dorsetmuscling × Merino (PDm × M), Merino × Merino (M × M) and Border Leicester × Merino (BL × M) genotypes were compared. Sires were selected on the basis of growth (PDg, M and BL) and muscle (PDm) using estimated breeding values (EBVs). Approximately 150 mixed sex animals were slaughtered at each age (595 in total for the four slaughters) and carcass quality traits, including hot carcass weight, loin and semitendinosus (ST) muscle weights, GR fat, fat C, rump fat depth and eye muscle area (EMA), were determined at 45 min to 3 days postslaughter. With age, all carcass quality traits increased in magnitude. Merinos produced the lightest and leanest carcasses (P < 0.01) with lowest values in nearly all the carcass quality traits. At 4 and 8 months of age carcass and loin weights, EMA and GR fat measures were greater (P < 0.01) in second-cross PDg × BLM sheep than all other genotypes. However, traits related to muscle content were similar between first- and second-cross Poll Dorset genotypes at 14 and 22 months of age. The BL × M genotype had lower values than the PD × M genotypes for these traits, at all ages. Traits related to carcass fatness showed that sheep with Border Leicester genetics tend to deposit more fat (P < 0.001) in the carcass than those with Merino or Poll Dorset genetics from 14 months of age onwards. Most carcass quality traits did not differ (P > 0.05) between first-cross Poll Dorset genotypes selected for growth or muscling (PDg × M v. PDm × M). Sires with high muscling EBVs are associated more with shorter carcass length than low muscling EBVs. From ~8 months of age, wethers had greater values (P < 0.05) than ewes of the same age for most carcass quality traits related to size and muscle development, but no effect was present for carcass quality traits that were related to fat deposition. It was clear that the carcass weight of M × M at 8 and 14 months were similar to that observed in PDg × BLM at 4 and 8 months, respectively. Loin weights of M × M and BL × M at 13 and 8 months were similar to that weights observed in PDg × BLM at 8 and 4 months of age, respectively. Animals slaughtered to a muscle weight specification will reach the target at a younger age when they have more Poll Dorset than Border Leicester background genetics. Alternatively animals slaughtered at a specified fat score (using fat C or rump fat depth) will reach the specification at a younger age when they have more Border Leicester than Poll Dorset genetics.

scholarly journals Genetics of meat quality and carcass traits and the impact of tenderstretching in two tropical beef genotypes

2009 ◽  
Vol 49 (6) ◽  
pp. 383 ◽  
Author(s):  
M. L. Wolcott ◽  
D. J. Johnston ◽  
S. A. Barwick ◽  
C. L. Iker ◽  
J. M. Thompson ◽  
...  
Keyword(s):  
Meat Quality ◽  
Shear Force ◽  
Carcass Traits ◽  
Intramuscular Fat ◽  
Quality Traits ◽  
Muscle Area ◽  
Live Animal ◽  
Eye Muscle ◽  
Meat Quality Traits ◽  
Lower Shear

Meat quality and carcass traits were measured for 2180 feedlot finished Brahman (BRAH) and Tropical Composite (TCOMP) steers to investigate genetic and non-genetic influences on shear force, and other meat quality traits. Genetic and phenotypic correlations were estimated between carcass and meat quality traits, and with live animal measurements collected in steers from weaning to feedlot exit, and their heifer half-sibs up to their first mating, which were managed in Australia’s tropical or subtropical environments. Left sides of carcasses were tenderstretched (hung by the aitch-bone) while right sides were conventionally hung (by the Achilles tendon). Tenderstretching reduced mean shear force by 1.04 kg, and phenotypic variance by 77% of that observed in conventionally hung sides. Genotype differences existed for carcass traits, with TCOMP carcasses significantly heavier, fatter, with greater eye muscle area, and lower retail beef yield than BRAH. TCOMP had lower shear force, and higher percent intramuscular fat. Meat quality and carcass traits were moderately heritable, with estimates for shear force and compression of 0.33 and 0.19 for BRAH and 0.32 and 0.20 for TCOMP respectively. In both genotypes, estimates of heritability for carcass traits (carcass weight, P8 and rib fat depths, eye muscle area and retail beef yield) were consistently moderate to high (0.21 to 0.56). Shear force and compression were genetically correlated with percent intramuscular fat (r g = –0.26 and –0.57, respectively), and meat colour (r g = –0.41 and –0.68, respectively). For TCOMP, lower shear force was genetically related to decreased carcass P8 fat depth (r g = 0.51). For BRAH steers and heifers measured at pasture, fatness traits and growth rates were genetically correlated with shear force, although the magnitude of these relationships varied with time of measurement. Net feed intake was significantly genetically correlated with carcass rib fat depth (r g = 0.49), eye muscle area (r g = –0.42) and retail beef yield (r g = –0.61). These results demonstrate that selection to improve production and carcass traits can impact meat quality traits in tropically adapted cattle, and that genotype specific evaluations will be necessary to accommodate different genetic relationships between meat quality, carcass and live animal traits.

scholarly journals Prediction of meat quality traits in the abattoir using portable near-infrared spectrometers: heritability of predicted traits and genetic correlations with laboratory-measured traits

2021 ◽  
Vol 12 (1) ◽  
Author(s):  
Simone Savoia ◽  
Andrea Albera ◽  
Alberto Brugiapaglia ◽  
Liliana Di Stasio ◽  
Alessio Cecchinato ◽  
...  
Keyword(s):  
Meat Quality ◽  
Near Infrared ◽  
External Validation ◽  
Genetic Correlations ◽  
Genetic Selection ◽  
Quality Traits ◽  
Eye Muscle ◽  
Meat Quality Traits ◽  
Color Traits ◽  
Selection Of

Abstract Background The possibility of assessing meat quality traits over the meat chain is strongly limited, especially in the context of selective breeding which requires a large number of phenotypes. The main objective of this study was to investigate the suitability of portable infrared spectrometers for phenotyping beef cattle aiming to genetically improving the quality of their meat. Meat quality traits (pH, color, water holding capacity, tenderness) were appraised on rib eye muscle samples of 1,327 Piemontese young bulls using traditional (i.e., reference/gold standard) laboratory analyses; the same traits were also predicted from spectra acquired at the abattoir on the intact muscle surface of the same animals 1 d after slaughtering. Genetic parameters were estimated for both laboratory measures of meat quality traits and their spectra-based predictions. Results The prediction performances of the calibration equations, assessed through external validation, were satisfactory for color traits (R2 from 0.52 to 0.80), low for pH and purge losses (R2 around 0.30), and very poor for cooking losses and tenderness (R2 below 0.20). Except for lightness and purge losses, the heritability estimates of most of the predicted traits were lower than those of the measured traits while the genetic correlations between measured and predicted traits were high (average value 0.81). Conclusions Results showed that NIRS predictions of color traits, pH, and purge losses could be used as indicator traits for the indirect genetic selection of the reference quality phenotypes. Results for cooking losses were less effective, while the NIR predictions of tenderness were affected by a relatively high uncertainty of estimate. Overall, genetic selection of some meat quality traits, whose direct phenotyping is difficult, can benefit of the application of infrared spectrometers technology.

scholarly journals Estimation of environmental effects and genetic parameters of carcass traits on Chikso (Korean brindle cattle)

2020 ◽  
Vol 33 (4) ◽  
pp. 525-530 ◽  
Author(s):  
Byoungho Park ◽  
Tae Jeong Choi ◽  
Mi Na Park ◽  
Sang-Hyon Oh
Keyword(s):  
Linear Model ◽  
Environmental Effects ◽  
Genetic Parameters ◽  
Coat Color ◽  
Carcass Traits ◽  
National Level ◽  
Pedigree Information ◽  
Small Scale ◽  
Muscle Area ◽  
Eye Muscle

Objective: The purpose of this study was i) to identify the characteristics of carcass traits in Chikso by gender, region, age at slaughter, and coat color using the carcass data collected from the nationwide pedigree information and coat color investigation, and ii) to estimate genetic parameters for breed improvement.Methods: A linear model was used to analyze the environmental effects on the carcass traits and to estimate genetic parameters. Analysis of variance was performed using TYPE III sum of squares for the unbalanced data provided by the general linear model procedure. Variance components for genetic parameters was estimated using REMLF90 of the BLUPF90 family programs.Results: Phenotypic performance of carcass weight (CW), eye muscle area (EMA), and backfat thickness (BF) in Chikso were lower than those of Hanwoo. This is a natural outcome because Hanwoo have undergone significant efforts for improvement at the national level, a phenomenon not observed in Chikso. Another factor influencing the above outcome was the smaller population size of Chikso compared to that of Hanwoo’s. The heritabilities of CW, EMA, BF, and marbling score in Chikso were estimated as 0.50, 0.37, 0.35, and 0.53, respectively, which were was higher than those of Hanwoo.Conclusion: Based on the genetic parameters that were estimated in this study, it is expected that the carcass traits will improve when the livestock research institutes at each province conduct small-scale performance tests and the semen is provided to farmers after selecting proven bulls using the state-of-art selection technique such as genomic selection.

scholarly journals Effect of boars own performance on progeny fattening and carcass traits in two different environments

2004 ◽  
Vol 20 (3-4) ◽  
pp. 55-63 ◽  
Author(s):  
I. Bahelka ◽  
P. Fľak ◽  
Anna Lukácová
Keyword(s):  
Performance Test ◽  
Average Daily Gain ◽  
Carcass Traits ◽  
Live Weight ◽  
Feed Consumption ◽  
Muscle Area ◽  
Eye Muscle ◽  
Carcass Parameters ◽  
Performance Traits ◽  
Progeny Performance

The effect of own performance traits of meat breed boars on fattening and carcass parameters of progeny in two different test stations (Bucany and Nitra) was evaluated. Own performance traits of boars were average daily gain (ADG) from birth to 100 kg live weight, backfat thickness (BF) and lean meat content (LMC) in field conditions. Progeny of boars was housed in pairs (gilt and barrow) and fed standardized feed mixture semi ad libitum. Progeny test lasted from 30 to 100 kg live weight. There were evaluated following parameters: ADG and feed consumption/kg gain (FC) at test from 30 to 100 kg live weight, slaughter weight (SW), proportion of meaty cuts (PMC) proportion of ham (PHAM), eye muscle area (EMA), and BF. At Bucany and Nitra was found the effect of genotype of boars on progeny BF and/or FC respectively. Better tested boars from own performance test individually as well as a group achieved in progeny better fattening and carcass traits than worse tested boars at Bucany (+33 g ADG, -0.21 cm BF, +2.34 % PMC). Progeny performance from better evaluated boars at Nitra did not exceed the progeny performance from worse tested boars. There was found significant effect of dams on progeny performance.

scholarly journals Integrating Single-Step GWAS and Bipartite Networks Reconstruction Provides Novel Insights into Yearling Weight and Carcass Traits in Hanwoo Beef Cattle

Animals ◽  
2020 ◽  
Vol 10 (10) ◽  
pp. 1836
Author(s):  
Masoumeh Naserkheil ◽  
Abolfazl Bahrami ◽  
Deukhwan Lee ◽  
Hossein Mehrban
Keyword(s):  
Beef Cattle ◽  
Complex Traits ◽  
Genome Wide Association Study ◽  
Additive Genetic Variance ◽  
Carcass Traits ◽  
Single Step ◽  
Bipartite Networks ◽  
Muscle Area ◽  
Eye Muscle ◽  
Network Analyses

In recent years, studies on the biological mechanisms underlying complex traits have been facilitated by innovations in high-throughput genotyping technology. We conducted a weighted single-step genome-wide association study (WssGWAS) to evaluate backfat thickness, carcass weight, eye muscle area, marbling score, and yearling weight in a cohort of 1540 Hanwoo beef cattle using BovineSNP50 BeadChip. The WssGWAS uncovered thirty-three genomic regions that explained more than 1% of the additive genetic variance, mostly located on chromosomes 6 and 14. Among the identified window regions, seven quantitative trait loci (QTL) had pleiotropic effects and twenty-six QTL were trait-specific. Significant pathways implicated in the measured traits through Gene Ontology (GO) term enrichment analysis included the following: lipid biosynthetic process, regulation of lipid metabolic process, transport or localization of lipid, regulation of growth, developmental growth, and multicellular organism growth. Integration of GWAS results of the studied traits with pathway and network analyses facilitated the exploration of the respective candidate genes involved in several biological functions, particularly lipid and growth metabolism. This study provides novel insight into the genetic bases underlying complex traits and could be useful in developing breeding schemes aimed at improving growth and carcass traits in Hanwoo beef cattle.

scholarly journals Genomic Analysis Using Bayesian Methods under Different Genotyping Platforms in Korean Duroc Pigs

Animals ◽  
2020 ◽  
Vol 10 (5) ◽  
pp. 752 ◽  
Author(s):  
Jungjae Lee ◽  
Yongmin Kim ◽  
Eunseok Cho ◽  
Kyuho Cho ◽  
Soojin Sa ◽  
...  
Keyword(s):  
Genomic Prediction ◽  
Genome Wide Association Study ◽  
Genomic Analysis ◽  
Snp Markers ◽  
Snp Genotyping ◽  
Muscle Area ◽  
Bayesian Approaches ◽  
Breeding Values ◽  
Estimated Breeding Values ◽  
Response Variables

Genomic evaluation has been widely applied to several species using commercial single nucleotide polymorphism (SNP) genotyping platforms. This study investigated the informative genomic regions and the efficiency of genomic prediction by using two Bayesian approaches (BayesB and BayesC) under two moderate-density SNP genotyping panels in Korean Duroc pigs. Growth and production records of 1026 individuals were genotyped using two medium-density, SNP genotyping platforms: Illumina60K and GeneSeek80K. These platforms consisted of 61,565 and 68,528 SNP markers, respectively. The deregressed estimated breeding values (DEBVs) derived from estimated breeding values (EBVs) and their reliabilities were taken as response variables. Two Bayesian approaches were implemented to perform the genome-wide association study (GWAS) and genomic prediction. Multiple significant regions for days to 90 kg (DAYS), lean muscle area (LMA), and lean percent (PCL) were detected. The most significant SNP marker, located near the MC4R gene, was detected using GeneSeek80K. Accuracy of genomic predictions was higher using the GeneSeek80K SNP panel for DAYS (Δ2%) and LMA (Δ2–3%) with two response variables, with no gains in accuracy by the Bayesian approaches in four growth and production-related traits. Genomic prediction is best derived from DEBVs including parental information as a response variable between two DEBVs regardless of the genotyping platform and the Bayesian method for genomic prediction accuracy in Korean Duroc pig breeding.

Indices of cellular development in muscles of lambs are influenced by sire estimated breeding values and pastoral nutritional system

2006 ◽  
Vol 57 (6) ◽  
pp. 651 ◽  
Author(s):  
P. L. Greenwood ◽  
G. E. Gardner ◽  
R. S. Hegarty
Keyword(s):  
Muscle Growth ◽  
Genetic Selection ◽  
Inverse Association ◽  
Semitendinosus Muscle ◽  
Eye Muscle ◽  
Weaning Weight ◽  
Breeding Values ◽  
Sire Group ◽  
Selection For ◽  
Estimated Breeding Values

This study examined influences of sire (n = 9) estimated breeding values (EBVs), sire-group (Muscle, Growth, and Control), and nutrition (low and high quality and availability pasture) from birth to slaughter at ~8 months of age on indices of muscle cellularity and transcriptional and translational capacity in 56 castrate lambs. Effects of nutritional systems to 8 months of age were greater, overall, than those due to EBVs or sire-group. Amount of DNA increased with increasing EBV for post-weaning eye muscle depth (PEMD or Muscle EBV) in longissimus but not in semimembranosus and semitendinosus muscles, while Muscle EBV also had an inverse association with concentration of DNA. Protein to DNA and RNA to DNA were related positively to Muscle EBV, the associations being strongest for the semitendinosus muscle. Post-weaning weight (PWWT or Growth) EBV correlated positively with the RNA to DNA ratio and, among high but not low nutrition lambs, was inversely related to concentration of muscle DNA, whereas post-weaning fat depth (PFAT or Fat) EBV was correlated positively with RNA concentration. Overall, the magnitude of effects of sire-group was less than for sire EBVs, presumably due to differing selection pressures for muscling, fatness, and growth. High nutrition lambs had more protein to DNA than low nutrition lambs in the longissimus and semitendinosus muscles, but not in the semimembranosus muscle. In low compared with high nutrition lambs, concentration of DNA was greater in the longissimus and semitendinosus muscles. Total amount of DNA was reduced by more in low compared with high nutrition in the longissimus and semimembranosus than in the semitendinosus, and amount of protein was reduced by more in low compared with high nutrition in the longissimus than in the other two muscles. We conclude that genetic selection for eye muscle depth in sheep has differing effects on cellular characteristics of the longissimus, semimembranosus, and semitendinosus muscles, and has greater effects on muscle cellular characteristics than genetic selection for post-weaning weight or fat depth.

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