Influence of species proportion and timing of establishment on stem quality in mixed red alder – Douglas-fir plantations

2004 ◽  
Vol 34 (4) ◽  
pp. 863-873 ◽  
Author(s):  
Amy T Grotta ◽  
Barbara L Gartner ◽  
Steven R Radosevich
Keyword(s):  
Stand Structure ◽  
Stand Density ◽  
Douglas Fir ◽  
Stem Form ◽  
Mixed Stands ◽  
Red Alder ◽  
Stem Quality ◽  
Multiple Stems ◽  
Old Trees ◽  
Species Proportion

The relationships among stand structure, Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) branch characteristics, and red alder (Alnus rubra (Bong.)) stem form attributes were explored for 10- to 15-year-old trees growing in mixed Douglas-fir – red alder plantations. Treatments included a range of species proportions, and red alder was either planted simultaneously with Douglas-fir or after 5 years. Both replacement effects (total stand density held constant) and additive effects (stand density doubled) of competition were considered. When the two species were planted simultaneously and red alder proportion was low, red alder trees had low crown bases and much stem defect (lean, sweep, and multiple stems). Douglas-fir grew slowly when the two species were planted simultaneously. When red alder planting was delayed, species proportion did not affect red alder stem form, and height to the base of the Douglas-fir live crown decreased with increasing red alder proportion. Doubling Douglas-fir density increased the height to the base of the Douglas-fir live crown; however, doubling stand density by adding red alder did not affect Douglas-fir crown height. Douglas-fir lumber coming from mixed stands may be inferior because of the changes in knot characteristics associated with these different patterns of crown recession. In stands with a low proportion of red alder, red alder product recovery may be compromised because of the stem defects described above.

Effects of species mixtures on growth and stand development of Douglas-fir and red alder

2006 ◽  
Vol 36 (3) ◽  
pp. 768-782 ◽  
Author(s):  
Steven R Radosevich ◽  
David E Hibbs ◽  
Claudio M Ghersa
Keyword(s):  
Pacific Northwest ◽  
Tree Mortality ◽  
Stand Structure ◽  
Stand Density ◽  
Douglas Fir ◽  
Forest Understory ◽  
Red Alder ◽  
Structure Tree ◽  
Species Proportions ◽  
Species Mixtures

In the Pacific Northwest, a mixture of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and red alder (Alnus rubra Bong.) often results when red alder regenerates naturally in planted conifer stands. The relationships among stand structure, tree mortality, tree size, and understory development in the two species mixtures were explored at two sites for the first 16 years after planting. Treatments included a range of species proportions, and red alder was either planted simultaneously with Douglas-fir or planting was delayed for 5 years. Red alder was also removed from some simultaneously planted proportions. Both replacement effects (total stand density held constant) and additive effects (stand density doubled) of the interaction were considered. Red alder grew relatively better at Cascade Head Experimental Forest in the Coast Range, while Douglas-fir grew better at H.J. Andrews Experimental Forest in the less temperate Cascade Mountains. Possible production benefits from mixed plantings were examined using two methods of calculation. Potential production benefits from certain planted proportions of the two species occurred at H.J. Andrews Experimental Forest. No planting time or species proportion resulted in yield improvements over monoculture stands at Cascade Head Experimental Forest. Understory species also varied because of differences in site and stand characteristics that resulted from the differences in planting times and species proportions.

Comparing productivity of pure and mixed Douglas-fir and western hemlock plantations in the Pacific Northwest

2006 ◽  
Vol 36 (6) ◽  
pp. 1484-1496 ◽  
Author(s):  
M M Amoroso ◽  
E C Turnblom
Keyword(s):  
Pacific Northwest ◽  
Stand Density ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Mixed Stand ◽  
Mixed Stands ◽  
Individual Tree ◽  
The Pacific ◽  
Increase In Productivity

We studied pure and 50/50 mixtures of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) plantations to compare attained total yields between mixed-species stands as opposed to monocultures of equal densities. Whether overall stand density influences this outcome has not been adequately investigated, and to address this we included three density levels (494, 1111, and 1729 trees/ha) in the analysis. At age 12, as components of the mixed stands, Douglas-fir exhibited greater height, diameter, and individual-tree volume than western hemlock at all densities. At 494 and 1111 trees/ha the monocultures had a higher volume per hectare than the mixed stand, but at 1729 trees/ha the mixed stand appeared to be just as productive as the pure stands. The increase in productivity by the mixture at high densities seems to have resulted from the partial stratification observed and most likely also from better use of the site resources. Because of this, less interspecific competition was probably experienced in the mixed stand than intraspecific competition in the pure stands. This study shows the important role density plays in the productivity of mixed stands and thus in comparing mixed and pure stands.

Spatial extent of impact of red alder on soil chemistry of adjacent conifer stands

1992 ◽  
Vol 22 (9) ◽  
pp. 1434-1437 ◽  
Author(s):  
Charles C. Rhoades ◽  
Dan Binkley
Keyword(s):  
Soil Ph ◽  
Soil Chemistry ◽  
Douglas Fir ◽  
Spatial Extent ◽  
N Availability ◽  
Soil N ◽  
Mixed Stands ◽  
Red Alder ◽  
Soil N Availability ◽  
River Site

We examined patterns in soil N availability and pH along transects extending from mixed stands of conifers (mostly Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco)) and red alder (Alnusrubra Bong.) to pure conifer stands at two locations. At the relatively infertile Wind River site, increased N availability was apparent for about 8–12 m downslope of the alder–conifer stand, but no effect was apparent upslope. At the fertile Cascade Head site, no trend was apparent in N availability across the stand boundaries, but soil pH in the conifer stand was depressed for about 5 m from the alder–conifer stand. Overall, the effects of alder on soil chemistry appeared limited to a distance of less than half the height of the trees.

Stand structure, thinning prescriptions, and density indexes in a Douglas-fir thinning study, Western Washington, U.S.A.

1983 ◽  
Vol 13 (1) ◽  
pp. 126-136 ◽  
Author(s):  
Chadwick Dearing Oliver ◽  
Marshall D. Murray
Keyword(s):  
Stand Structure ◽  
Volume Growth ◽  
Basal Area ◽  
Stand Density ◽  
Close Relation ◽  
Douglas Fir ◽  
Volume Number ◽  
Large Trees ◽  
Before And After ◽  
Number Of Stems

A Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) thinning study was established in 1959 in a stand begun after logging in 1930. Thinnings to set basal area densities were done in 1959, 1962, 1966, and 1970. On each plot both large and small trees were removed since average basal area per tree was kept constant before and after thinning. Volume growth varied greatly between plots of the same age, initial basal area, and site because of differences in stand structure. Large trees on a plot grew more per tree and per basal area than small trees. Stand basal area, stand volume, number of stems, or number of dominant and codominant trees were not closely related to volume growth per hectare, although density indexes giving weight to larger trees showed the closest relation. The lack of close relation between stand density indexes and growth found here and elsewhere probably means the indexes do not uniquely define stand structures; it does not necessarily mean that thinning will not increase volume growth per hectare. Volume growth per hectare after thinning to a given basal area density will be greater and probably more consistent if larger trees are left and enough time is allowed for the stand to recover following thinning.

Effects of stand density on the growth of young Douglas-fir trees

2002 ◽  
Vol 32 (3) ◽  
pp. 420-427 ◽  
Author(s):  
David R Woodruff ◽  
Barbara J Bond ◽  
Gary A Ritchie ◽  
William Scott
Keyword(s):  
Growth Response ◽  
Stand Density ◽  
Early Growth ◽  
Douglas Fir ◽  
Volume Index ◽  
Annual Rings ◽  
Wood Cellulose ◽  
Positive Growth ◽  
Second Year ◽  
Old Trees

The objectives of this study were (i) to provide further evidence of a positive correlation of stand density with early growth of coastal Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii); (ii) to determine when after planting the positive growth response occurs and how long it lasts; and (iii) to use stable isotopes of carbon to test whether the mechanism(s) responsible for the positive growth response to density are related to variables affecting photosynthesis, such as nutrient or moisture availability. We measured annual height (h) and diameter (d) growth (retrospectively) of 8- and 12-year-old trees in initial planting densities of 300, 1360, and 2960 trees/ha. Both height and diameter growth increased with density through the fifth year after planting and decreased with density by year 7. Diameter squared × height (d2h) was used as a volume index to assess increase in tree volume. Second-year increase in d2h for the high-density treatments was 300% of that in the low-density treatments. The δ13C values of wood cellulose from annual rings of the second and third years after planting were not significantly different among densities, suggesting either (i) no significant differences in the effects of water availability, nutrient availability, or source air on photosynthesis in the three density treatments or (ii) differences that produced no net effect on δ13C.

Size of Douglas-fir trees in relation to distance from a mixed red alder–Douglas-fir stand

1993 ◽  
Vol 23 (11) ◽  
pp. 2413-2418 ◽  
Author(s):  
Richard E. Miller ◽  
Donald L. Reukema ◽  
Timothy A. Max
Keyword(s):  
Stand Density ◽  
Positive Influence ◽  
Poor Quality ◽  
Douglas Fir ◽  
Mixed Stand ◽  
Stem Volume ◽  
Nitrogen Fixing ◽  
Red Alder ◽  
Wide Strip ◽  
Southwest Washington

Variation in diameter, height, and stem volume of 57-year-old Douglas-fir (Pseudotsugamenziesii var. menziesii (Mirb.) Franco) was related to distance of these trees from a 27 m wide strip in the same Douglas-fir plantation that had been interplanted with red alder (Alnusrubra Bong.). Within the interplanted strip and despite its greater total stand density, bole volume of dominant and codominant Douglas-fir averaged 1.27 m3 compared with 0.55, 0.45, 0.46, or 0.49 m3 in trees 15, 30,45, or 60 m, respectively, from the edge of the mixed stand. Some positive influence of nitrogen-fixing red alder apparently extended about 15 m beyond the edge of the mixed stand at this poor quality site in southwest Washington. We infer that similar ribbonlike distributions of naturally regenerated red alder could be retained to improve growth of nearby conifers on nitrogen-deficient sites.

Stand structure and growing space efficiency following thinning in an even-aged Douglas-fir stand

1988 ◽  
Vol 18 (7) ◽  
pp. 859-866 ◽  
Author(s):  
K. L. O'Hara
Keyword(s):  
Stand Structure ◽  
Volume Growth ◽  
Basal Area ◽  
Stand Density ◽  
Douglas Fir ◽  
Stem Volume ◽  
Projection Area ◽  
Individual Tree ◽  
Space Efficiency ◽  
Individual Trees

The growth of individual trees from four thinning treatments in a 64-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stand was analyzed to determine desirable residual stand structures after thinning. Dominant and codominant trees had the highest individual tree stem volume growth rates over the previous 5 years, and accounted for most stand volume growth in thinned and unthinned stands. Two measures of growing space, crown projection area and sapwood basal area (a surrogate for leaf area), were used to measure how efficiently individual trees used their growing space. Crown classes were useful in characterizing growing space efficiency (volume growth per unit of growing space) only in the unthinned treatment. In thinned treatments, tall trees with medium-sized crowns were most efficient, while in the unthinned treatment, tall trees with relatively large crowns were most efficient. A large crown in an unthinned stand was comparable in size to a medium-sized crown in a thinned stand. Results suggest growing space is not limiting individual tree growth in thinned stands and that thinning to a particular stand structure is more appropriate than thinning to a particular level of stand density.

scholarly journals Low Thinning as a Forest Restoration Tool at Redwood National Park

2011 ◽  
Vol 26 (2) ◽  
pp. 91-93 ◽  
Author(s):  
Jason R. Teraoka ◽  
Christopher R. Keyes
Keyword(s):  
Forest Restoration ◽  
Stand Structure ◽  
Basal Area ◽  
Stand Density ◽  
Variable Density ◽  
Douglas Fir ◽  
Redwood National Park ◽  
Second Growth ◽  
Need To Evaluate

Abstract A growing interest in the restoration of young second-growth forests by managers of reserves in the redwood region has led to a need to evaluate restoration-based silvicultural strategies. This case study assessed the effectiveness of low thinning as a forest restoration tool via analysis of stand structure at Redwood National Park's Whiskey Forty Forest Restoration Study. The second-growth stand had more than 5,500 trees ha−1 and 57.0 m2 ha−1 basal area and consisted chiefly of three species: Douglas-fir (the dominant species), redwood, and tanoak. Low thinning reduced stand density but also reduced species richness by eliminating scarce species. Seven years after thinning, growth was enhanced (33.6% gain in basal area), and mortality was minor (3% of all stems); however, Douglas-fir remained competitive in the upper canopy. Its average basal area increment was less than redwood's, but its radial growth was equal and its rate of basal area growth was greater in the years following thinning. We conclude that the thinning improved stand conditions but did not fully satisfy restoration goals and that other thinning methods, such as variable-density thinning, are likely to be more effective at promoting redwood dominance.

scholarly journals Structural analysis of mixed stands coming from natural regeneration and plantations after fire

Web Ecology ◽  
2010 ◽  
Vol 10 (1) ◽  
pp. 32-37
Author(s):  
T. Tsitsoni ◽  
M. Tsakaldimi ◽  
E. Simeliadou ◽  
M. Fouska
Keyword(s):  
Stand Structure ◽  
Structural Characteristics ◽  
Basal Area ◽  
Stem Diameter ◽  
Diameter Distribution ◽  
Cupressus Sempervirens ◽  
Mixed Stands ◽  
Crown Length ◽  
Stem Quality ◽  
Naturally Regenerated Stands

Abstract. We analyzed the structure and growth of naturally regenerated stands of Pinus brutia that mixed with planted broad-leaved and conifer species, 12 years after wildfire and examined the degree of species mix. Field data on stand structure of P. brutia forest were taken in spring 2009 on northern and southern aspects differing in regeneration conditions. Sixteen sample plots were selected and all individuals and their attributes measured. The results showed that in northern aspects the forest is composed of P. brutia in the over-storey and Quercus pubescens and Cupressus sempervirens in the under-storey, while in southern aspects the forest was mainly composed by P. brutia (81%). Stem diameter distribution of P. brutia in both aspects followed almost a normal pattern. All P. brutia individuals were characterized by vigorous growth and good to normal stem quality. Aspect did not statistically affect structural characteristics of P. brutia trees and saplings. However, on the northern aspect stem diameter, height, crown length and basal area of P. brutia were greater than in the southern aspect. Aspect significantly affected structural characteristics of Q. pubescens.

ROOT SPREAD CAN BE ESTIMATED FROM CROWN WIDTH OF DOUGLAS FIR, LODGEPOLE PINE, AND OTHER BRITISH COLUMBIA TREE SPECIES

1964 ◽  
Vol 40 (4) ◽  
pp. 456-473 ◽  
Author(s):  
J. Harry G. Smith
Keyword(s):  
Lodgepole Pine ◽  
Stand Density ◽  
Douglas Fir ◽  
Red Alder ◽  
Rooting Zone ◽  
Red Cedar ◽  
Crown Width ◽  
Maximum Root ◽  
Poorly Drained Soils ◽  
Root Spread

Knowledge of the relationships between lateral root spread and crown width is needed to guide plans for providing optimum space per tree during silvicultural operations. Crown width is a valuable indicator of root spread of open-grown Douglas fir, lodgepole and ponderosa pines, and white, Engelmann and Sitka spruces. The association between root spread and crown width is less reliable in forest-grown trees but still useful.Detailed studies of root systems were made by excavation of roots of trees blown down in the U.B.C. Campus Forest by a typhoon in October, 1962. Roots of 89 Douglas fir, 81 western hemlock, 61 western red cedar, and 33 red alder trees were mapped and analyzed in relation to 18 tree and stand variables. In addition, average and maximum root depth, number of main roots, the ratio of maximum to average root spread, and portion of rooting zone occupied by roots were studied.Average and maximum radius of roots of Douglas fir and lodgepole pine were analyzed in relation to crown width, d.b.h., height, age and other tree and stand characteristics. Appropriate regression and correlation analyses were completed for all species and the possible advantages of ratio estimates were noted.Ratios of root spread to crown width were influenced by species, stand density, and kind of soil. Ratios averaged 1.1 for open- and 0.9 for forest-grown Douglas fir but were 2.4 for both open- and forest-grown lodgepole pine on peat or poorly drained soils.

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