scholarly journals Interspecific variation in bristle number on forewings of tiny insects does not influence clap-and-fling aerodynamics

Author(s):  
Vishwa T. Kasoju ◽  
Daniel S. Moen ◽  
Mitchell P. Ford ◽  
Truc T. Ngo ◽  
Arvind Santhanakrishnan

Miniature insects must overcome significant viscous resistance in order to fly. They typically possess wings with long bristles on the fringes and use clap-and-fling mechanism to augment lift. These unique solutions to the extreme conditions of flight at tiny sizes (< 2 mm body length) suggest that natural selection has optimized wing design for better aerodynamic performance. However, species vary in wingspan, number of bristles (n), and bristle gap (G) to diameter (D) ratio (G/D). How this variation relates to body length (BL) and its effects on aerodynamics remain unknown. We measured forewing images of 38 species of thrips and 21 species of fairyflies. Our phylogenetic comparative analyses showed that n and wingspan scaled positively and similarly with body length across both groups, whereas G/D decreased with BL, with a sharper decline in thrips. We next measured aerodynamic forces and visualized flow on physical models of bristled wings performing clap-and-fling kinematics at chord-based Reynolds number of 10 using a dynamically scaled robotic platform. We examined the effects of dimensional (G, D, wingspan) and non-dimensional (n, G/D) geometric variables on dimensionless lift and drag. We found that: (a) increasing G reduced drag more than decreasing D; (b) changing n had minimal impact on lift generation; and (c) varying G/D minimally affected aerodynamic forces. These aerodynamic results suggest little pressure to functionally optimize n and G/D. Combined with the scaling relationships between wing variables and BL, much wing variation in tiny flying insects might be best explained by underlying shared growth factors.

2020 ◽  
Author(s):  
Vishwa T. Kasoju ◽  
Mitchell P. Ford ◽  
Truc T. Ngo ◽  
Arvind Santhanakrishnan

ABSTRACTFlight-capable miniature insects of body length (BL) < 2 mm typically possess wings with long bristles on the fringes. Though their flight is challenged by needing to overcome significant viscous resistance at chord-based Reynolds number (Rec) on the order of 10, these insects use clap-and-fling mechanism coupled with bristled wings for lift augmentation and drag reduction. However, inter-species variation in the number of bristles (n) and inter-bristle gap (G) to bristle diameter (D) ratio (G/D) and their effects on clap-and-fling aerodynamics remain unknown. Forewing image analyses of 16 species of thrips and 21 species of fairyflies showed that n and maximum wing span were both positively correlated with BL. We conducted aerodynamic force measurements and flow visualization on simplified physical models of bristled wing pairs that were prescribed to execute clap-and-fling kinematics at Rec=10 using a dynamically scaled robotic platform. 23 bristled wing pairs were tested to examine the isolated effects of changing dimensional (G, D, span) and non-dimensional (n, G/D) geometric variables on dimensionless lift and drag. Within biologically observed ranges of n and G/D, we found that: (a) increasing G provided more drag reduction than decreasing D; (b) changing n had minimal impact on lift generation; and (c) varying G/D produced minimal changes in aerodynamic forces. Taken together with the broad variation in n (32-161) across the species considered here, the lack of impact of changing n on lift generation suggests that tiny insects may experience reduced biological pressure to functionally optimize n for a given wing span.SUMMARY STATEMENTIntegrating morphological analysis of bristled wings seen in miniature insects with physical model experiments, we find that aerodynamic forces are unaffected across the broad biological variation in number of bristles.


2014 ◽  
Vol 118 (1201) ◽  
pp. 229-266 ◽  
Author(s):  
S. N. Nguyen ◽  
E. S. Greenhalgh ◽  
J. M. R. Graham ◽  
A. Francis ◽  
R. Olsson

AbstractLarge transport aircraft are particularly susceptible to impact damage from runway debris thrown up by the landing gear. A methodology was developed to predict the trajectories of stones lofted by the nose wheel and subjected to aerodynamic forces due to the wake behind the nose landing gear and beneath the aircraft. In conjunction with finite element modelling of the stone/ground/tyre contact mechanics, an analytical model was used to perform a stochastic prediction of the trajectories of runway stones to generate impact threat maps which showed the relative likelihood of stones impinging upon various areas on the underside of a C-130 Hercules. The impact envelopes for the C-130 extended three to eighteen metres behind the nose wheel and two metres either side of the centre of the aircraft. The impact threat maps were especially sensitive to the values of the coefficients of lift and drag acting on the stone during its flight.


Author(s):  
Haibin Xuan ◽  
Jun Hu ◽  
Yong Yu ◽  
Jiaolong Zhang

Recently, numerous studies have been conducted to clarify the effects of corrugation wing on aerodynamic performances. The effects of the corrugation patterns and inclination angles were investigated using computational fluid dynamic method in gliding and hovering flight at Reynolds numbers of order 104. The instantaneous aerodynamic forces and the vorticity field around the wing models were provided to research the underlying mechanisms of aerodynamic effects of corrugated wing models. The findings can be concluded as follows: (1) the corrugation patterns have different effects on aerodynamic performance. The effect of noncamber corrugated wing is to decrease the lift and increase drag compared with a flat-plate when the angle of attack is less than 25° during gliding flight. The corrugated wing with a camber (corrug-2) after the valleys enhances the aerodynamic forces when angle of attack is higher than 35°. The valley inclination angle has limited effect on aerodynamic forces in gliding flight. (2) The lift forces of different corrugation patterns show significantly asymmetric during the upstroke and downstroke. The main reason leads to this phenomenon is the case that two sides of the corrugated wings are not symmetric around the pitching axis. The corrugated wing with only two valleys (corrug-1) changes the lift and drag very slightly. Corrug-2 produces larger peak during downstroke and smaller peak during upstroke. The increase in the inclination angle has limited effect on the aerodynamic forces. The possible reason for these small aerodynamic effects might be that the corrugated wings are smoothed by small vortices trapped in valleys. The main reason for the significant difference between plate and corrug-2 is that the recirculating vortices trapped in the saddle and hump reduce the pressure above the wing surface.


A proper understanding of how locusts fly must be based upon knowledge of how the wings are moved. A desert locust was suspended from a balance and placed in an air stream so that it flew under nearly the same conditions as during natural forward flight. Four stroboscopic slow-motion films were selected for measurement. The movements of the wings, i.e. their positions, velocities and accelerations, were then calculated in sufficient detail to show how these quantities vary with time during one complete wing stroke. The aerodynamic lift and drag of the entire natural wing were measured in a wind tunnel with the wing arranged in different positions relative to the flow. By placing it in the boundary layer of the tunnel, the wind speed was graded from tip to base in approximately the same way as during the actual flight. There is therefore no error due to scale effect or to the induced drag. In most respects the wings resemble ordinary, slightly cambered airfoils. Their characteristics are given as polar diagrams. The kinematic and aerodynamic analyses make it possible to calculate the forces which act upon the locust at any instant of time. It is here necessary to presuppose that the non-stationary flight situations are essentially similar to a sequence of stationary situations. For locusts, this presupposition is justified: (i) from theoretical estimates of the quantitative effect of non-stationary flow; and (ii) from control measurements of the average thrust and lift produced during flight. It was found that the calculated vertical force, when averaged over an entire wing stroke, equalled the average reduction in body weight, as measured directly on the flight balance. Similarly, the average thrust of the wings corresponded to the drag of the body. The analysis shows how the aerodynamic forces vary during the wing stroke. The hindwings are responsible for about 70 % of the total lift and thrust. About 80 % of the lift is produced during the downstroke. During flight at normal lift the angles of attack (middle part of wing) are small during the upstroke and vary between 10 and 15° during the downstroke. When the lift was larger or smaller than the body weight these figures increased or decreased respectively. The forewings are peculiar in two ways: (i) during the middle part of the downstroke a true flap (the vannus) is put into action; (ii) during the upstroke the proximal part has a Z-shaped cross-section and gives but little lift and drag. The hindwings are characteristic in that the posterior part (vannus) is flexible and becomes moulded by the wind, increasing the angle of attack at which stalling occurs to about 25°. Since both the movements of the wings relative to the body and the aerodynamic forces are known at any instant, the exchange of power with the surrounding air can be calculated. The moments of inertia of the wing mass being known, the power for accelerating the wings can also be estimated. The sum of these contributions is the power which passes the wing fulcrum; this estimate is used in a later paper (part IX) where the energetics of flight is discussed in detail. The diagrams are correct to scale. The restriction of freedom caused by the suspension is discussed, together with the possible errors of a stationary analysis.


eLife ◽  
2021 ◽  
Vol 10 ◽  
Author(s):  
Ben J Hightower ◽  
Patrick W A Wijnings ◽  
Rick Scholte ◽  
Rivers Ingersoll ◽  
Diana D Chin ◽  
...  

How hummingbirds hum is not fully understood, but its biophysical origin is encoded in the acoustic nearfield. Hence, we studied six freely hovering Anna's hummingbirds, performing acoustic nearfield holography using a 2176 microphone array in vivo, while also directly measuring the 3D aerodynamic forces using a new aerodynamic force platform. We corroborate the acoustic measurements by developing an idealized acoustic model that integrates the aerodynamic forces with wing kinematics, which shows how the timbre of the hummingbird's hum arises from the oscillating lift and drag forces on each wing. Comparing birds and insects, we find that the characteristic humming timbre and radiated power of their flapping wings originates from the higher harmonics in the aerodynamic forces that support their bodyweight. Our model analysis across insects and birds shows that allometric deviation makes larger birds quieter and elongated flies louder, while also clarifying complex bioacoustic behavior.


AVIA ◽  
2021 ◽  
Vol 2 (2) ◽  
Author(s):  
Y Parlindungan ◽  
S Tobing

This study is inspired by the flapping motion of natural flyers: insects. Many insects have two pairs of wings referred as tandem wings. Literature review indicates that the effects of tandem wing are influenced by parameters such as stagger (the stream-wise distance between the aerodynamic center of the front and the rear airfoil), angle-of-attack and flow velocity. As a first stage, this study focuses on the effects of stagger (St) on the aerodynamic performance of tandem wings. A recent numerical study of stagger on tandem airfoils in turbulent flow (Re = 6000000) concluded that a larger stagger resulted in a decrease in lift force, and an increase in drag force. However, for laminar flow (Re = 2000), increasing the stagger was not found to be detrimental for aerodynamic performance. Another work also revealed that the maximum lift coefficient for a tandem configuration decreased with increasing stagger. The focus of this study is to perform an experimental analysis of tandem two-dimensional (2D) NACA 0012 airfoils. The two airfoils are set at the same angle-of-attack of 0° to 15° with 5° interval and three variations of stagger: 1c, 1.5c and 2c. The experiments are conducted using an open-loop-subsonic wind tunnel at a Reynolds number of 170000. The effects of St on the aerodynamic forces (lift and drag) are analyzed


2013 ◽  
Author(s):  
Dennis Evangelista ◽  
Griselda Cardona ◽  
Eric Guenther-Gleason ◽  
Tony Huynh ◽  
Austin Kwong ◽  
...  

We report the effects of posture and morphology on the static aerodynamic stability and control effectiveness of physical models based on the feathered dinosaur,Microraptor gui, from the Cretaceous of China. Postures had similar lift and drag coefficients and were broadly similar when simplified metrics of gliding were considered, but they exhibited different stability characteristics depending on the position of the legs and the presence of feathers on the legs and the tail. Both stability and the function of appendages in generating maneuvering forces and torques changed as the glide angle or angle of attack were changed. These are significant because they represent an aerial environment that may have shifted during the evolution of directed aerial descent and other aerial behaviors. Certain movements were particularly effective (symmetric movements of the wings and tail in pitch, asymmetric wing movements, some tail movements). Other appendages altered their function from creating yaws at high angle of attack to rolls at low angle of attack, or reversed their function entirely. WhileM. guilived afterArchaeopteryxand likely represents a side experiment with feathered morphology, the general patterns of stability and control effectiveness suggested from the manipulations of forelimb, hindlimb and tail morphology here may help understand the evolution of flight control aerodynamics in vertebrates. Though these results rest on a single specimen, as further fossils with different morphologies tested, the findings here could be applied in a phylogenetic context to reveal biomechanical constraints on extinct flyers arising from the need to maneuver. Now published in PLOS ONE http://dx.plos.org/10.1371/journal.pone.0085203


2012 ◽  
Vol 79 (2) ◽  
Author(s):  
A. J. Vance ◽  
J. M. Buick ◽  
J. Livesey

This paper describes the aerodynamic forces on a rugby ball traveling at speeds between 5 and 15 ms−1. This range is typical of the ball speed during passing play and a range of kicking events during a game of rugby, and complements existing data for higher velocities. At the highest speeds considered here, the lift and drag coefficients are found to be compatible with previous studies at higher velocities. In contrast to these higher speed investigations, a significant variation is observed in the aerodynamic force over the range of velocities considered. Flow visualizations are also presented, indicating how the flow pattern, which is responsible for the aerodynamic forces, changes with the yaw angle of the ball. This flow and, in particular, the position of the separation points, is examined in detail. The angular position of the separation point is found to vary in a linear manner over much of the surface of the rugby ball; however, this behavior is interrupted when the separation point is close to the ‘tip’ of the ball.


1996 ◽  
Vol 316 ◽  
pp. 173-196 ◽  
Author(s):  
Chien-Cheng Chang ◽  
Sheng-Yuan Lei

The present study aims at relating lift and drag to flow structures around a delta wing of elliptic section. Aerodynamic forces are analysed in terms of fluid elements of non-zero vorticity and density gradient. The flow regime considered is Mα = 0.6 ∼ 1.8 and α = 5° ∼ 19°, where Mα denotes the free-stream Mach number and α the angle of attack. Let ρ denote the density, u velocity, and ω vorticity. It is found that there are two major source elements Re(x) and Ve(x) which contribute about 95% or even more to the aerodynamic forces for all the cases under consideration, \[R_e({\bm x})=-\frac{1}{2} {\bm u}^2 \nabla\rho \cdot \nabla\phi\quad {\rm and}\quad V_e ({\bm x}) = -\rho{\bm u}\times {\bm \omega}\cdot \nabla\phi,\] where θ is an acyclic potential, generated by the delta wing moving with unit velocity in the negative direction of the force (lift or drag). All the physical quantities are non-dimensionalized. Detailed force contributions are analysed in terms of the flow structures and the elements Re(x) and Ve(x). The source elements Re(x) and Ve(x) are concentrated in the following regions: the boundary layer in front of (below) the delta wing, the primary and secondary vortices over the delta wing, and a region of expansion around the leading edge. It is shown that Ve(x) due to vorticity prevails as the source of forces at relatively low Mach number, Mα < 0.7. Above about Mα = 0.75, Re(x) due to compressibility generally becomes the dominating contributor to the lift, while the overall contribution from Ve(x) decreases with increasing Mα, and even becomes negative at Mα = 1.2 for the lift, and at a higher Mα for the drag. The analysis is carried out with the aid of detailed numerical results by solving the Reynolds-averaged Navier–Stokes equations, which are in close agreement with experiments in comparisons of the surface pressure distributions.


2020 ◽  
Vol 306 ◽  
pp. 05005
Author(s):  
Haibin Xuan ◽  
Jun Hu ◽  
Yong Yu ◽  
Jiaolong Zhang

Some insects possess corrugated wings, which distinguish from the ordinary airfoils. It is important to research the corrugation effect on the aerodynamic performances. A series of corrugated wing models were designed based on former research in represent study to find out the underlying mechanisms. The effects of the corrugation pattern and inclination angle were studied using computational fluid dynamic (CFD) method during hovering flight at Reynolds numbers in the order of 104. The instantaneous aerodynamic forces and the vorticity field around the wing models were provided. The findings are as follows: (1) the results of this paper show that patterns of corrugation have different effect on aerodynamic performances. The corrugated wing like Corrug-1 changes the lift and drag very slightly compared with a flat-plate. The corrugation patterns like Corrug-2 and 3 of wing model reduce the lift and drag force. (2) the increase in the inclination angle has limited effect the aerodynamic forces. The inclination angles like corrug-3 and 4 produce almost the same forces.


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