The Study of a Predator-Prey Model with Fear Effect Based on State-Dependent Harvesting Strategy

In presence of predator population, the prey population may significantly change their behavior. Fear for predator population enhances the survival probability of prey population, and it can greatly reduce the reproduction of prey population. In this study, we propose a predator-prey fishery model introducing the cost of fear into prey reproduction with Holling type-II functional response and prey-dependent harvesting and investigate the global dynamics of the proposed model. For the system without harvest, it is shown that the level of fear may alter the stability of the positive equilibrium, and an expression of fear critical level is characterized. For the harvest system, the existence of the semitrivial order-1 periodic solution and positive order- q ( q ≥ 1 ) periodic solution is discussed by the construction of a Poincaré map on the phase set, and the threshold conditions are given, which can not only transform state-dependent harvesting into a cycle one but also provide a possibility to determine the harvest frequency. In addition, to ensure a certain robustness of the adopted harvest policy, the threshold condition for the stability of the order- q periodic solution is given. Meanwhile, to achieve a good economic profit, an optimization problem is formulated and the optimum harvest level is obtained. Mathematical findings have been validated in numerical simulation by MATLAB. Different effects of different harvest levels and different fear levels have been demonstrated by depicting figures in numerical simulation using MATLAB.

Bifurcation analysis of the predator–prey model with the Allee effect in the predator

The use of predator–prey models in theoretical ecology has a long history, and the model equations have largely evolved since the original Lotka–Volterra system towards more realistic descriptions of the processes of predation, reproduction and mortality. One important aspect is the recognition of the fact that the growth of a population can be subject to an Allee effect, where the per capita growth rate increases with the population density. Including an Allee effect has been shown to fundamentally change predator–prey dynamics and strongly impact species persistence, but previous studies mostly focused on scenarios of an Allee effect in the prey population. Here we explore a predator–prey model with an ecologically important case of the Allee effect in the predator population where it occurs in the numerical response of predator without affecting its functional response. Biologically, this can result from various scenarios such as a lack of mating partners, sperm limitation and cooperative breeding mechanisms, among others. Unlike previous studies, we consider here a generic mathematical formulation of the Allee effect without specifying a concrete parameterisation of the functional form, and analyse the possible local bifurcations in the system. Further, we explore the global bifurcation structure of the model and its possible dynamical regimes for three different concrete parameterisations of the Allee effect. The model possesses a complex bifurcation structure: there can be multiple coexistence states including two stable limit cycles. Inclusion of the Allee effect in the predator generally has a destabilising effect on the coexistence equilibrium. We also show that regardless of the parametrisation of the Allee effect, enrichment of the environment will eventually result in extinction of the predator population.

EFFECT OF ADDITIONAL FOOD ON PREDATOR–PREY INTERACTIONS WITH WATER-LEVEL FLUCTUATION

Significant variations of the water-level of the lake can have a strong impact on the persistence of species. Indeed, when the water-level is low, during the autumn, the contact between the predator and the prey is more frequent, and the predation increases. Conversely, when the water-level is high, in the spring, it is more difficult for the predator to find a prey and the predation decreases. In this paper, we consider a seasonally varying predator–prey model to study the influence of water-level variations on the interaction between two species of fishes in an artificial lake. A seasonal variation of the water-level is introduced in the predation rate. The predator population is provided some additional food apart from the focal prey, and follows logistic growth in the absence of prey population. As control upon the over predation, the predator population is harvested. Sensitivity analysis shows that the biomass of predator population is highly sensitive to the additional food and water variations. In the absence of additional food, our results show bursting patterns of fishes whereas positive periodic solution arises if the additional food is available in sufficient amount. The positive periodic solution is shown to be globally stable. Higher values of water-level fluctuations induce double periodic oscillations. Our findings show that providing additional food source to the generalist predator together with water-level fluctuations exerts a strong influence on the interaction between fishes.

Evolutionary rescue can prevent rate-induced tipping

The transformation of ecosystems proceeds at unprecedented rates. Recent studies suggest that high rates of environmental change can cause rate-induced tipping. In ecological models, the associated rate-induced critical transition manifests during transient dynamics in which populations drop to dangerously low densities. In this work, we study how indirect evolutionary rescue—due to the rapid evolution of a predator’s trait—can save a prey population from the rate-induced collapse. Therefore, we explicitly include the time-dependent dynamics of environmental change and evolutionary adaptation in an eco-evolutionary system. We then examine how fast the evolutionary adaptation needs to be to counteract the response to environmental degradation and express this relationship by means of a critical rate. Based on this critical rate, we conclude that indirect evolutionary rescue is more probable if the predator population possesses a high genetic variation and, simultaneously, the environmental change is slow. Hence, our results strongly emphasize that the maintenance of biodiversity requires a deceleration of the anthropogenic degradation of natural habitats.

Stability and Hopf Bifurcation of a Stage-Structured Cannibalism Model with Two Delays

In this work, we consider a stage-structured cannibalism model with two delays. One delay characterizes the lag effect of negative feedback of the prey species, the other has the effect of gestation of the adult predator population. Firstly, criteria for the local stability of feasible equilibria are established. Meanwhile, by choosing delay as a bifurcation parameter, the criteria on the existence of Hopf bifurcation are established. Furthermore, by the normal form theory and center manifold theorem, we derive the explicit formulas determining the properties of periodic solutions. Finally, the theoretical results are illustrated by numerical simulations, from which we can see that the predator’s gestation time delay can make the chaotic phenomenon disappear and maintain periodic oscillation, and that a large feedback time delay of prey can make predators extinct and prey form a periodic solution.

Contrasting effects of prey refuge on biodiversity of species

Refugia have been perceived as a major role in structuring species biodiversity, and understanding the impacts of this force in a community assembly with prey–predator species is a difficult task because refuge process can interact with different ecological components and may show counterintuitive effects. To understand this problem, we used a simple two-species model incorporating a functional response inspired by a Holling type-II equation and a prey refuge mechanism that depends on prey and predator population densities (i.e., density-dependent prey refuge). We then perform the co-dimension one and co-dimension two bifurcation analysis to examine steady states and its stability, together with the bifurcation points as different parameters change. As the capacity of prey refuge is varied, there occur critical values i.e., saddle-node and supercritical Hopf bifurcations. The interaction between these two co-dimension one bifurcations engenders distinct outcomes of ecological system such as coexistence of species, bistability phenomena and oscillatory dynamics. Additionally, we construct a parameter space diagram illustrating the dynamics of species interactions as prey refuge intensity and predation pressure vary; as the two saddle-node move nearer to one another, these bifurcations annihilate tangentially in a co-dimension two cusp bifurcation. We also realised several contrasting observations of refuge process on species biodiversity: for instance, while it is believed that some refuge processes (e.g., constant proportion of prey refuge) would result in exclusion of predator species, our findings show that density-dependent prey refuge is beneficial for both predator and prey species, and consequently, promotes the maintenance of species biodiversity.

Age-selective harvesting in a delayed predator–prey model with fear effect

In this article, we discussed the dynamic behavior of a delay-induced harvested predator–prey model with fear effects (perceived by the prey). We then considered selective harvesting terms for both species which provide some fixed elapsed time to the prey and for the predator population before they are harvested. In other words, we are limiting the harvesting of species below a certain age so that they can grow to a certain specific size or age and thus protect juvenile populations. Reproduction of the prey population can also be greatly impeded due to the influence of the fear effect. The consideration of selective harvesting together with the effect of fear on the proposed system to show stable coexistence to the oscillatory mode and vice versa via Hopf-bifurcation. For better ecological management of the community, our study reveals the fact that collection delays and intensities should be maintained. Numerical simulations were performed to validate our analytical results.

Aplikasi Pestisida Nabati Maja - Gadung dan Metabolit Sekunder Beauveria Bassiana Bals. Untuk Mengendalikan Hama Thrips Sp. Pada Tanaman Cabai Rawit

Pest attacks on chili plants are the main limiting factor in cultivation activities. The research aims to determine the effect of single application of botanical pesticide maja-gadung (PMG), single application of secondary metabolite Beauveria bassiana BIO (BIO B10), combined application of PMG and BIO B10 on pest populations, predator populations and plant growth and production of chili. The reesearh used a factorial Completely Randomized Block Design (RAKL) with 3 replications, so there were 9 treatment combinations (0 ml/l PMG, 4 ml/l PMG, 8 ml/l BIO B10, 0 ml/l PMG, 2 ml/l BIO B10, 4 ml/l BIO B10, 4 ml PMG+2 ml/l BIO B10, 4 ml/ml PMG+4 ml BIO B10, 8 ml/l PMG+2ml/l BIO B10, 8 ml/l PMG+4 ml/l BIO B10). The variables observed were Thrips sp population, predator population, plant height, number of leaves, and fruit weight per plant. The results showed that the single application treatment of PMG with a concentration of 4 ml/l and 8 ml/l was able to suppress the population of Thrips sp. by 21.6% and 41.4% compared to the control. The single application treatment of BIO B10 with concentrations of 2 ml/l and 4 ml/l was able to suppress the population of Thrips sp. by 66.5% and 65.5% compared to the control. The best combination application is PMG and BIO B10with a concentration of 8 ml/l + 4 ml/l which is able to suppress the population of Thrips sp. by 76.9% compared to the control. The application treatment of PMG andBIO B10 did not affect plant height, number of leaves, and fruit weight per plant.