Changes in root morphology after precommercial thinning in balsam fir stands

2003 ◽  
Vol 33 (12) ◽  
pp. 2452-2459 ◽  
Author(s):  
Jean-Claude Ruel ◽  
Catherine Larouche ◽  
Alexis Achim
Keyword(s):  
Radial Growth ◽  
Abies Balsamea ◽  
Stand Density ◽  
Balsam Fir ◽  
Cross Sectional ◽  
Beam Shape ◽  
Precommercial Thinning ◽  
Beneficial Effects ◽  
Density Control ◽  
Over Time

Precommercial thinning is often used to control stand density in naturally regenerated balsam fir (Abies balsamea (L.) Mill.) stands. Early stand density control could have beneficial effects on longer term stand stability through a modification of stem shape and root development. To assess the effect of precommercial thinning, two thinned and two unthinned stands were selected. Root sections were collected at 25 cm from the centre of the stem for all major roots (diameter greater than 2 cm). A disk was also cut at breast height level. From these disks, response in radial growth was determined. Detailed root measurements were taken on the sample disks to assess treatment effect on the following variables: asymmetry in root cross-sectional area (CSA) distribution, individual root shape, as well as changes in root shape over time. A quick and pronounced response in root growth occurred. This response was greater than that observed in the trunk. No asymmetry in root CSA distribution was observed at the stand level. Trees allocated more to radial growth above the biological centre of the root both in treated and control stands, but this trend was increased by thinning. Roots also tended to develop T-beam shapes over time, both in control and thinned stands. Most roots initially did not possess an I-beam shape and did not develop one during the course of the study. According to our results, trees respond quickly to the new growing conditions created by thinning by increasing biomass allocation to parts of the roots where mechanical stresses are greater.

Long-term outcome of precommercial thinning in northwestern New Brunswick: growth and yield of balsam fir and red spruce

2008 ◽  
Vol 38 (3) ◽  
pp. 592-610 ◽  
Author(s):  
Doug Pitt ◽  
Len Lanteigne
Keyword(s):  
Block Design ◽  
Abies Balsamea ◽  
Red Spruce ◽  
Balsam Fir ◽  
Growth And Yield ◽  
Stem Volume ◽  
Long Term Outcome ◽  
Precommercial Thinning ◽  
Minimum Requirement

A study was established between 1959 and 1961 to study the long-term responses of balsam fir ( Abies balsamea (L.) Mill.) and red spruce ( Picea rubens Sarg.) to precommercial thinning. Three nominal spacings of 4 ft (1.2 m), 6 ft (1.8 m), and 8 ft (2.4 m) were compared with an unthinned control in a randomized complete block design with five replicates. At the time of thinning, natural regeneration averaged 16 years of age, 8 years after harvest. Although thinning had minimal effect on gross total volume production over a 42 to 44 year observation period, actual spacings between 2.1 and 2.5 m produced an average of 360 m3·ha–1gross merchantable volume (GMV), representing a 21% gain over unthinned stands. The same spacings produced quadratic mean diameters of 21 and 23 cm, respectively, compared with 18 cm in the unthinned stands. These size increases translated to individual stem volume gains of 33% and 62%, significantly reducing the age at which thinned stands would meet a specified minimum requirement for merchantability or habitat. The mean annual increment of GMV ranged from 6 m3·ha–1·year–1in unthinned stands, to more than 7 m3·ha·–1·year–1in the thinned stands, and had not yet culminated an average of 50 years postharvest.

Effects of precommercial thinning on the forest value chain in northwestern New Brunswick: Part 4 – Lumber production, quality and value

2013 ◽  
Vol 89 (04) ◽  
pp. 474-489 ◽  
Author(s):  
Isabelle Duchesne ◽  
Doug G. Pitt ◽  
Francis Tanguay
Keyword(s):  
Value Chain ◽  
Positive Impact ◽  
Abies Balsamea ◽  
Basic Density ◽  
Production Quality ◽  
Balsam Fir ◽  
Appreciable Effect ◽  
Precommercial Thinning ◽  
Product Value ◽  
Negative Effect

Forty-eight years after application, three nominal precommercial thinning (PCT) spacings of 4 ft (1.2 m), 6 ft (1.8 m) and 8 ft (2.4 m) were compared to an unthinned control in six replicate, balsam fir (Abies balsamea [L.] Mill.)-dominated stands. Within three of these replicates chosen for clearcut harvest in 2008, a total of 150 trees were bucked into 479 sawlogs that produced 10 829 board feet of lumber. PCT had a positive impact on Premium lumber grade recovery per tree, increasing it from an average of 9 foot board measure (fbm) in the unthinned control plots, to 22 fbm in plots nominally spaced to 8 ft (p = 0.02). No. 2 & Better grade yields increased from 42 to 76 fbm/tree in these same sets of plots (p < 0.01). PCT to 6 ft and 8 ft increased the production of 2×4 and 2×6 lumber over the control (p ≤ 0.04). Stand-level gross total product value increased between 11% and 23% in thinned stands (estimated at +$3000 to +$6600 per ha, p < 0.01). PCT had no appreciable effect on sawn lumber wood density (p ≥ 0.26) or lumber stiffness (MOE; ≤-6.2%, p ≥ 0.11), but had a negative effect on lumber strength (MOR; ≤ -13.4%, p ≤ 0.03) and wood basic density at stump height (≤- 7%; p < 0.01). MDF panel properties were generally good, regardless of spacing. We recommend thinning young balsam fir stands to a maximum nominal spacing of 6 ft (1.2 m) to maintain satisfactory lumber mechanical properties and maximize product value per hectare.

AN ANALYSIS OF MEASURES OF DENSITY IN EVEN-AGED BALSAM FIR AND JACK PINE STANDS

1964 ◽  
Vol 40 (4) ◽  
pp. 474-481 ◽  
Author(s):  
P. E. Vezina
Keyword(s):  
Pinus Banksiana ◽  
Abies Balsamea ◽  
Basal Area ◽  
Stand Density ◽  
Jack Pine ◽  
Balsam Fir ◽  
Aerial Photographs ◽  
Growing Stock ◽  
The One ◽  
Crown Closure

The concept of stand density in relation to thinning is examined and its development over the years is discussed. Present difficulties of objectively measuring stand density are recognized and probable future trends towards the development of better formulae to express stand density are outlined. Researchers should continue to collect information on interrelationships among stand variables. Certain merits accrue from description of stand density in terms of variables, such as crown closure, that can be measured with some precision from aerial photographs. Conversely, valid estimates of crown closure which are often difficult to obtain by means of devices from the ground, could be predicted from stand density. Three stand variables, used as expressions of stand density, were tested in crown closure simple regressions in even-aged, unmanaged stands of balsam fir (Abies balsamea (L.) Mill.) and jack pine (Pinus banksiana Lamb.). These are: total number of trees, number of trees 4 inches and up, and basal area per acre. The strongest relationship found was the one where crown closure is compared with basal area; it was stronger for jack pine than for balsam fir. This was explained by differences in tolerance among the two species. The significance of these relationships for the stand development, and the feasibility of using height-and diameter-based indices as measures of growing stock in studies of yield are discussed.

Missing growth rings at the trunk base in suppressed balsam fir saplings

2002 ◽  
Vol 32 (10) ◽  
pp. 1776-1783 ◽  
Author(s):  
Sylvain Parent ◽  
Hubert Morin ◽  
Christian Messier
Keyword(s):  
Radial Growth ◽  
Adventitious Roots ◽  
Abies Balsamea ◽  
Dry Mass ◽  
Balsam Fir ◽  
Leaf Biomass ◽  
Growth Rings ◽  
Terminal Bud ◽  
Below Ground ◽  
Age Estimates

Numerous researchers have suggested a causal relationship between low leaf biomass in suppressed trees and the lack of radial growth at the base of the trunk. The objective of this study was to verify this relationship with suppressed balsam fir (Abies balsamea (L.) Mill.) saplings found growing in an old-growth fir stand. A total of 29 saplings varying in height from 67 to 183 cm were uprooted. All saplings had adventitious roots. All terminal bud scars (TBS) found between the apex of the terminal leader and the trunk base (first adventitious root) as well as those found below ground were localized, and rings were counted between TBS along the aboveground trunk. Various morphological traits and the ratio of photosynthetic tissue dry mass (P, needles) to non-photosynthetic tissue dry mass (nP, aboveground stem) were used as an indicator of tree vigour. Between 3 and 33 rings counted along the aboveground trunk were missing at the trunk base. The number of missing rings at the base of the trunk was correlated with total height (r = 0.41), height growth (r = –0.51), radial growth (r = –0.44), the P/nP ratio (r = –0.73), and the proportion of live crown (r = –0.62). Moreover, from 2 to 35 additional rings, missing at the trunk base, were found in the belowground section of trunk and these missing rings were associated with the adventitious roots phenomenon. In conclusion, suppressed firs had missing rings at the base of the trunk. When all of the missing rings were added to the number of rings counted at the base of the trunk, age estimates provided a different temporal pattern of recruitment compared with that obtained by solely counting rings at the base of the trunk. Stem analysis on the entire trunk is the best aging method for suppressed balsam fir saplings.

scholarly journals Lower Plasma Total Testosterone Levels Were Associated With Steeper Decline in Brain Glucose Metabolism in Non-demented Older Men

2021 ◽  
Vol 13 ◽  
Author(s):  
Xiwu Wang ◽  
Zhaoting Lv ◽  
Qian Wu ◽  
Huitao Liu ◽  
Yanrou Gu ◽  
...  
Keyword(s):  
Glucose Metabolism ◽  
Older People ◽  
Total Testosterone ◽  
Cross Sectional ◽  
Brain Glucose ◽  
Brain Glucose Metabolism ◽  
Testosterone Levels ◽  
Beneficial Effects ◽  
Significant Difference ◽  
Over Time

ObjectiveThere is growing evidence that testosterone may be implicated in the pathogenesis of Alzheimer’s disease (AD). We aimed to examine the relationship between plasma total testosterone levels and change in brain glucose metabolism over time among non-demented older people.MethodsThe association of plasma total testosterone levels with change in brain glucose metabolism among non-demented older people was investigated cross-sectionally and longitudinally. Given a significant difference in levels of plasma total testosterone between gender, we performed our analysis in a sex-stratified way. At baseline, 228 non-demented older people were included: 152 males and 76 females.ResultsIn the cross-sectional analysis, no significant relationship between plasma total testosterone levels and brain glucose metabolism was found in males or females. In the longitudinal analysis, we found a significant association of plasma total testosterone levels with change in brain glucose metabolism over time in males, but not in females. More specifically, in males, higher levels of total testosterone in plasma at baseline were associated with slower decline in brain glucose metabolism.ConclusionWe found that higher levels of total testosterone in plasma at baseline were associated with slower decline in brain glucose metabolism in males without dementia, indicating that testosterone may have beneficial effects on brain function.

scholarly journals EFFECTS OF DEFOLIATION BY SPRUCE BUDWORM (CHORISTONEURA FUMIFERANA CLEM.) ON RADIAL GROWTH AT BREAST HEIGHT OF BALSAM FIR (ABIES BALSAMEA (L.) MILL.) AND WHITE SPRUCE (PICEA GLAUCA (MOENCH) VOSS.)

1958 ◽  
Vol 34 (1) ◽  
pp. 39-47 ◽  
Author(s):  
J. R. Blais
Keyword(s):  
Picea Glauca ◽  
Radial Growth ◽  
Choristoneura Fumiferana ◽  
Abies Balsamea ◽  
Spruce Budworm ◽  
White Spruce ◽  
Balsam Fir ◽  
First Year ◽  
Growth Ratio ◽  
Breast Height

The relationship between spruce budworm defoliation and radial growth at breast height for balsam fir and white spruce trees of merchantable size was studied in various stands in northwestern Ontario. Defoliation was recorded yearly for these stands from the beginning of the infestation, and radial growth measurements were obtained from increment cores. The first year of radial growth suppression was calculated by comparing the growth of the affected species with that of jack pine and red pine trees by means of a growth-ratio technique. Apparent suppression in balsam fir and white spruce varied between stands, and, generally, occurred at the earliest in the second year and at the latest in the fourth year of severe defoliation. A wide ring at the base of the tree coinciding with the first year of suppression as reported by Craighead was non-existent.

A stand density management diagram for balsam fir in New Brunswick

2006 ◽  
Vol 82 (5) ◽  
pp. 700-711 ◽  
Author(s):  
M. Penner ◽  
D E Swift ◽  
R. Gagnon ◽  
J. Brissette
Keyword(s):  
New Brunswick ◽  
Abies Balsamea ◽  
Stand Density ◽  
Maximum Size ◽  
Balsam Fir ◽  
Quadratic Mean ◽  
Stand Management ◽  
Forest Region ◽  
Quadratic Mean Diameter ◽  
Stand Density Management

A stand management density diagram (SDMD) is presented for balsam fir (Abies balsamea (L.) Mill.) forests in New Brunswick. The SDMD incorporates a maximum size density line, as well as quadratic mean diameter and top height isolines. Several mortality functions are evaluated. The resultant SDMD should be a useful tool for projecting early stand development and determining the timing and intensity of thinnings. Key words: Acadian Forest Region, mortality curves

Effects of precommercial thinning on the forest value chain in northwestern New Brunswick: Part 3 – Incidence of root and butt decay

2013 ◽  
Vol 89 (04) ◽  
pp. 464-473 ◽  
Author(s):  
Gary Warren ◽  
Patricia Baines ◽  
Jean Plamondon ◽  
Doug G. Pitt
Keyword(s):  
Forest Management ◽  
Value Chain ◽  
Abies Balsamea ◽  
Balsam Fir ◽  
Stand Age ◽  
Stem Form ◽  
Green River ◽  
Precommercial Thinning ◽  
Permanent Sample Plots ◽  
Butt Rot

The Green River precommercial thinning (PCT) trials were established between 1959 and 1961 in naturally regenerating balsam fir (Abies balsamea [L.] Mill.)-dominated stands an average of eight years after overstory removal. Three nominal spacings of 4 ft (1.2 m), 6 ft (1.8 m) and 8 ft (2.4 m) were compared to an unthinned control in six replicate blocks. In the fall of 2008, following completion of the ninth sequential evaluation of the study’s 48 permanent sample plots, three of the six replicates were clearcut harvested; butt rot data were collected immediately afterwards. To date, forest management research and goals have focused on the benefits of PCT, such as increased tree size and merchantable volume, shorter rotation ages, and better stem form and uniformity. Comparatively little attention has been placed on negative aspects of PCT, such as the incidence and development of root and butt rots, and their impact on fibre yields and wood product values. Results from the Green River study provide evidence that PCT may increase the incidence of butt rot in balsam fir, with incidence proportional to thinning intensity (p < 0.01). We also observed incidence and volume of butt rot to increase with stem diameter (p ≤ 0.05). The experiment suggests that factors such as stand age at time of thinning, and age at the time of harvest are important considerations when it comes to mitigating the impacts of butt rot through forest management.

Insect defoliators as major disturbance factors in the high-altitude balsam fir forest of Mount Mégantic, southern Quebec

1998 ◽  
Vol 28 (12) ◽  
pp. 1832-1842 ◽  
Author(s):  
Louise Filion ◽  
Serge Payette ◽  
Ann Delwaide ◽  
Najat Bhiry
Keyword(s):  
North America ◽  
High Altitude ◽  
Tree Ring ◽  
Radial Growth ◽  
Abies Balsamea ◽  
Spruce Budworm ◽  
Balsam Fir ◽  
Cumulative Impact ◽  
Tree Ring Data ◽  
Paper Birch

Tree-ring data from a mature balsam fir forest, located at the top of Mount Mégantic (elevation 1100 m), southern Quebec, suggest that insect defoliators were major disturbance factors in the development of high-altitude balsam fir forests. A comparison between the radial growth trend of balsam fir (Abies balsamea (L.) Mill.), a host species of the spruce budworm (Choristoneura fumiferana Clem.), and paper birch (Betula papyrifera Marsh), a nonhost species, showed that several growth depressions in the balsam fir chronology corrresponded to documented spruce budworm outbreaks in southern Quebec in the 1910s, 1950s, 1970s, and possibly in the 1870s. Tree mortality was extensive during the last infestation because of the relatively old age (>60 years) of many balsam fir and, possibly, to the cumulative impact of defoliation. The tree-ring series from paper birch showed several drops in radial growth after the 1930s, possibly related to the large-scale birch dieback that occurred in eastern North America. Macrofossil data (insect remains) from one sample of the uppermost organic soil layers (F horizon) confirm the presence of the spruce budworm at the study site. The ecological role of insect defoliators is discussed in the context of the high-altitude balsam fir forests in northeastern North America where abiotic disturbances are considered the primary controlling factors in stand dynamics.

Regeneration and stand development following a spruce budworm outbreak, spruce budworm inspired harvest, and salvage harvest

2012 ◽  
Vol 42 (10) ◽  
pp. 1759-1770 ◽  
Author(s):  
C. Eiry Spence ◽  
David A. MacLean
Keyword(s):  
Species Composition ◽  
Structural Characteristics ◽  
Choristoneura Fumiferana ◽  
Abies Balsamea ◽  
Canopy Cover ◽  
Spruce Budworm ◽  
Balsam Fir ◽  
Stand Development ◽  
Eastern Canada ◽  
Precommercial Thinning

Spruce budworm ( Choristoneura fumiferana (Clemens)) (SBW) outbreaks are a major disturbance that influence stand dynamics and succession in balsam fir ( Abies balsamea (L.) Mill.) dominated forests of eastern Canada. We used stand and regeneration data collected from five plots in each of one immature and five mature stands before, during, and 30 years following the last major SBW outbreak in the Cape Breton Highlands to examine the role of stand and regeneration characteristics in shaping future stand development. Comparisons were also made between regeneration in four SBW outbreak stands versus two SBW salvage stands, with and without a subsequent precommercial thinning, and with 25 plots that underwent SBW emulation harvest. In mature unharvested balsam fir stands, species composition 30 years following the SBW outbreak was closely related to predisturbance species composition, and in immature fir stands, hardwood composition increased from 0% to 4%–27%. Species composition in harvested stands varied depending on whether intolerant hardwoods had been precommercially thinned, where thinned stands had 30% less hardwood 30 years postdisturbance than unthinned stands. Seedling density decreased by 17%–85% from 1979 to 1989 in all SBW outbreak stands, but average seedling height increased by 17%–500% as live canopy cover decreased from an average of 50% to 4%. Results suggest that advanced regeneration should be protected during harvest of balsam fir dominated stands, post-outbreak precommercial thinning will increase tree growth, and live tree retention can help develop late-seral structural characteristics in second-growth stands.

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