Revision of Mesojassus Tillyard, 1916, from the Late Triassic of Queensland (Hemiptera: Cicadomorpha: Membracoidea: Archijassidae)

New specimens and a re-examination of their holotypes have clarified the status of six nominal species of the extinct membracoid family Archijassidae from the Late Triassic (Norian) fossil insect localities at Mount Crosby, Denmark Hill and Dinmore in south-eastern Queensland. The 57 available tegmina ostensibly attributable to one or other of the six species are remarkably similar in venation and only vary in size and to a lesser extent in shape. The latter character varies subtlety across a continuum and is of no use in species definition. The tegmina, however, fall into two distinct size groups, and in the absence of any other discernible or consistent diagnostic characters, these groups are adopted as separate species, acknowledging, of course, the artificial nature of fossil insect species based on the size only of isolated wings. The following taxonomic changes result: Mesojassus Tillyard, 1916 (= Triassojassus Tillyard, 1919, syn. nov., = Triassocotis Evans, 1956, syn. nov., = Hylicellites Becker-Migdisova, 1962, syn. nov.), Mesojassus ipsviciensis Tillyard, 1916 (= Triassojassus proavitus Tillyard, 1919, syn. nov., = Triassocotis stricta Evans, 1961, syn. nov.), Mesojassus australis (Evans, 1956) comb. nov. (= Triassocotis amplicata Evans, 1961, syn. nov., = Hylicellites reducta (Evans, 1956), syn. nov.). Mesojassus is one of the four genera of the subfamily Archija` ssinae, and differs from the Jurassic Archijassus Handlirsch, 1906, Mesoledra Evans, 1956, and Ardela Ansorge, 1996, in the separation of R and M well before the arculus (at the same level or slightly beyond in the others), the proximal position of dSc, well before the apex of the clavus (at or beyond the apex in the others), and the two-branched RA (simple in the others). The separation of R and M distinctly basal to the arculus, a character of frequent occurrence in extant membracoids, is proposed as a possible apomorphy for this otherwise most plesiomorphic genus of the Membracoidea. Mesojassus, the oldest member of the extant Membracoidea, is one of a growing inventory of genera from the Late Triassic of Queensland which are the oldest representatives of extant groups, adding further evidence of the Triassic as the dawn of much of the modern insect fauna.

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Revision of Mesothymbris Evans, 1956, from the Late Triassic of Mount Crosby, Queensland (Hemiptera: Cicadomorpha: Hylicelloidea: Hylicellidae)

Zootaxa ◽

10.11646/zootaxa.4629.3.6 ◽

2019 ◽

Vol 4629 (3) ◽

pp. 389-396

Author(s):

KEVIN J. LAMBKIN

Keyword(s):

Basal Cell ◽

The Other ◽

Late Triassic ◽

Nominal Species ◽

South Eastern ◽

Incertae Sedis ◽

Generic Composition ◽

The Status

New specimens and a re-examination of their holotypes have clarified the status of five species of the extinct cicadomorphan family Hylicellidae from the Late Triassic (Norian) Mount Crosby Formation of south-eastern Queensland. All were found to be conspecific, resulting in the following synonymies: Mesothymbris Evans, 1956 (= Triassoscytina Evans, 1956, syn. nov. = (in part) Triassoscytinopsis Evans, 1956, syn. nov.), Mesothymbris perkinsi Evans, 1956 (= Mesothymbris woodwardi Evans, 1956, syn. nov. = Triassoscytina incompleta Evans, 1956, syn. nov. = Triassoscytinopsis stenulata Evans, 1956, syn. nov. = Triassoscytinopsis aberrans Evans, 1956, syn. nov.). The Hylicellidae is still poorly defined as is its generic composition. Mesothymbris, however, is clearly distinct from the other Mount Crosby hylicellids, Hylicella Evans, 1956, and Triassocotis, Evans, 1956, in the quite distal primary fork of R, the angled RA at the point of separation of RA1 with RA2 directed towards the apex of the tegmen, the upright RA1, the shape of the intra-medial cell, and CuA just distal to the basal cell strongly curved and very closely approximating the claval suture. The new synonymies further clarify the composition of the cicadomorphan fauna of the Mount Crosby Formation, which as a result of this and other recent revisions, now comprises 16 nominal species in the Dysmorphoptilidae, Hylicellidae and Mesojabloniidae, as well as three species incertae sedis. In the presence of Dysmorphoptilidae and Hylicellidae, the Mount Crosby cicadomorphan fauna is similar to that of the younger Late Triassic Blackstone Formation at nearby Denmark Hill and Dinmore. It differs significantly, however, in the absence, after 90 years of collecting of 100s of specimens, of any representatives of the Dunstaniidae, Mesogereonidae, or Tettigarctidae, families so characteristic of the Denmark Hill/Dinmore fauna. Whether this difference is biogeographical, ecological, or simply as a result of differential preservation is unknown.

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III.—The Miocene Insect-fauna of Œningen, Baden

Geological Magazine ◽

10.1017/s0016756800005963 ◽

1895 ◽

Vol 2 (3) ◽

pp. 116-122 ◽

Cited By ~ 6

Author(s):

Samuel H. Scudder

Keyword(s):

Fossil Insects ◽

Nominal Species ◽

Insect Fauna ◽

Number Of Species ◽

Fossil Insect ◽

The World ◽

Comparative Zoology ◽

Louis Agassiz

Thanks solely to the labours of the late Oswald Heer, the fossil insect-fauna of Œningen is better known than that of any other locality or horizon in the world. But it is by no means so well known as it should be; for although Heer, in his latest enumeration of the specimens seen by him (Urwelt der Schweiz, 2e Aufl. 1879, p. 383), repeats precisely the same figures he has already given in 1861 (Recherches sur le climat du pays tert., p. 197), indicating in an interval of eighteen years no addition to his material (over 5000 specimens), his repeated additions to the number of species from that locality show that he had not fully worked over what he had. Indeed, thirty years ago, I arranged for exhibition in the Museum of Comparative Zoology at Cambridge a collection of nearly one hundred and fifty named species secured by Prof. Louis Agassiz from Heer, of which more than forty still remain undescribed; there are also a considerable number of merely nominal species enumerated by Heer in his Urwelt der Schweiz and elsewhere, duly catalogued by me in my Index to Fossil Insects (Bull. U. S. Geol. Surv. No. 71), but as yet neither described nor figured. In addition to this it may be noted that in the enumeration referred to above Heer mentions 543 species of beetles, while less than 270 nominal species have yet been published from Œningen, and only seven species from the Swiss Miocene, included in the enumeration.

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Recognition and partial solution of nomenclatural issues involving copepods of the family Monstrillidae (Crustacea: Copepoda: Monstrilloida)

Zootaxa ◽

10.11646/zootaxa.4486.4.5 ◽

2018 ◽

Vol 4486 (4) ◽

pp. 497 ◽

Cited By ~ 1

Author(s):

MARK J. GRYGIER ◽

EDUARDO SUÁREZ-MORALES

Keyword(s):

Original Description ◽

Partial Solution ◽

International Commission ◽

Zoological Nomenclature ◽

Nominal Species ◽

Separate Species ◽

Language Version ◽

Type Series ◽

The Status ◽

Registration Number

This work seeks to expose and clear up nomenclatural irregularities involving copepods of the order Monstrilloida, family Monstrillidae. The diagnostic text related to Monstrilla minuta Isaac, 1974 and four nominal species of Thaumaleus Krøyer, 1849 (now Cymbasoma Thompson, 1888) proposed by Isaac in 1974 is sufficient for all names to be available from their original description except for Thaumaleus similirostratus, which was proposed conditionally in 1974 and was first made available by Isaac in 1975; “similirostris” as used by Grygier in 1995 is an incorrect subsequent spelling. Four other specific names proposed in 1975 by Isaac, but disclaimed by him as nomina nuda (an action permitted retroactively by the Fourth Edition of the International Code of Zoological Nomenclature) have never been made available. By quoting the necessary information from Isaac’s doctoral dissertation, two of them are validated herein under the names Thaumaleus frondipes Isaac in Grygier & Suárez-Morales, sp. nov., and Strilloma scotti Isaac in Grygier & Suárez-Morales, sp. nov., and are immediately reassigned as new combinations to Cymbasoma and Monstrilla Dana, 1849, respectively. A fifth such name, Thaumaleus tumorifrons, has already been made available under the authorship of Suárez-Morales, 1999, but its females are excluded from the type series; the spelling of the specific name of the new species recently proposed for those females, Cymbasoma mediterranea Suárez-Morales, Goruppi, Olazabal & Tirelli, 2017, is emended to mediterraneum to match the gender of the genus. For Cymbasoma bowmani Suárez-Morales & Gasca, 1998, the “Form B” female mentioned in the original description is excluded from the type series. The authorship and date of availability of Haemocera (currently Cymbasoma) morii depends on which language version of Article 13.1.1 of the Code is followed; a ruling by the International Commission on Zoological Nomenclature under Article 87 of the Code is necessary to resolve the matter. The composition of the type series of Cymbasoma bullatum (Scott, 1909) in terms of both number and sex has become unclear; its type locality is restricted herein to the vicinity of Obi Island in the Moluccas. Despite a published statement to the contrary, the syntype series of Cymbasoma germanicum (Timm, 1893) included specimens from other localities than just Helgoland. The type series of Cymbasoma guerrerense Suárez-Morales & Morales-Ramírez, 2009 consists only of the holotype, which was mistakenly reported under the wrong registration number. The supposed invalidity of Monstrilla capitellicola Hartman, 1961 is discussed. Monstrilla javensis Isaac, 1974, nomen nudum, has remained unavailable owing to lack of adherence to Article 16.1 of the Code by later authors; the specific name is made available herein, under Suárez-Morales’ authorship, in the combination Cymbasoma javense sp. nov. The taxonomic (and eventual nomenclatural) question of the status of M. mariaeugeniae Suárez-Morales & Islas-Landeros, 1993 vis à vis M. wandelii Stephensen, 1913, i.e. as a separate species or a subspecies of the latter, remains unsettled. Cymbasoma lenticula Suárez-Morales & McKinnon, 2014 and Monstrillopsis boonwurrungorum Suárez-Morales & McKinnon, 2014 are fixed herein as the correct original spellings of those two specific names. Resolution of the problem posed by assignment of the specific name reticulata to supposedly non-conspecific males and females in the genus Monstrillopsis Sars, 1921 requires the designation of a neotype by the International Commission on Zoological Nomenclature.

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Taxonomic Uncertainty and the Anomaly Zone: Phylogenomics Disentangle a Rapid Radiation to Resolve Contentious Species (Gila robusta complex) in the Colorado River

10.1101/692509 ◽

2019 ◽

Cited By ~ 2

Author(s):

Tyler K. Chafin ◽

Marlis R. Douglas ◽

Max R. Bangs ◽

Bradley T. Martin ◽

Steven M. Mussmann ◽

...

Keyword(s):

Colorado River ◽

Nominal Species ◽

Rapid Radiation ◽

Evolutionary Mechanisms ◽

Phylogenetic Discordance ◽

The Status ◽

Taxonomic Uncertainty ◽

Morphological Differences ◽

Phylogenetic Models ◽

Methodological Difficulties

AbstractSpecies is an indisputable unit for biodiversity conservation, yet their delimitation is fraught with both conceptual and methodological difficulties. A classic example is the taxonomic controversy surrounding the Gila robusta complex in the lower Colorado River of southwestern North America. Nominal species designations were originally defined according to weakly diagnostic morphological differences that conflicted with traditional genetic analyses. Consequently, the complex was re-defined as a single polytypic unit, with the proposed ‘threatened’ status of two being withdrawn at the federal level. Here, we utilized dense spatial and genomic sampling (N=387 and >22k loci) to re-evaluate the status of the complex, based on SNP-based coalescent and polymorphism-aware phylogenetic models. In doing so, all three species were supported as evolutionarily independent lineages, despite widespread phylogenetic discordance. To understand this discrepancy with past studies, we categorized evolutionary mechanisms driving discordance. We tested (and subsequently rejected) prior hypotheses suggesting that phylogenetic discord in the complex was hybridization-driven. Instead, we found the G. robusta complex to have diverged within the ‘anomaly zone’ of tree space and, as such, have accumulated inconsistent patterns of diversity which have confounded prior studies. After extending these analyses with phylogeographic modeling, we propose that this is reflective of a rapid radiation promoted by Plio-Pleistocene tectonism. Our results not only support resurrection of the three species as distinct entities, but also offer an empirical example of how phylogenetic discordance can be categorized in other recalcitrant taxa where variation is primarily partitioned at the species-level.

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Mesozoic Coleopteran Faunas from Argentina: Geological Context, Diversity, Taphonomic Observations, and Comparison with Other Fossil Insect Records

Psyche A Journal of Entomology ◽

10.1155/2012/242563 ◽

2012 ◽

Vol 2012 ◽

pp. 1-14 ◽

Cited By ~ 7

Author(s):

María Belén Lara ◽

Oscar Florencio Gallego ◽

Lara Vaz Tassi

Keyword(s):

Late Jurassic ◽

Middle Triassic ◽

Late Triassic ◽

Animal Kingdom ◽

Fossil Insect ◽

San Luis ◽

Fundamental Information ◽

The World ◽

Geological Units ◽

Deseado Massif

The order Coleoptera is the most diversified group of the Class Insecta and is the largest group of the Animal Kingdom. This contribution reviews the Mesozoic insects and especially the coleopteran records from Argentina, based on bibliographical and unpublished materials (86 described species, 526 collected specimens). The material came from different geological units from the late Middle Triassic to the Late Triassic (Bermejo, Cuyo, and Malargüe basins) to the Middle-Late Jurassic and Early Cretaceous (Deseado Massif, Cañadón Asfalto, and San Luís Basin). The coleopteran record is composed of 29 described species with 262 collected specimens (isolated elytra) mainly represented by Triassic species and only four specimens recorded in Jurassic units, all of them currently unpublished. These fossil coleopterans provide fundamental information about the evolution of insects in the Southern Hemisphere and confirm the Triassic Argentinean insect deposits to be among the most important in the world.

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Taxonomic affinities and paleogeography of Stromatomorpha californica Smith, a distinctive Upper Triassic reef-adapted demosponge

Journal of Paleontology ◽

10.1666/08-146.1 ◽

2009 ◽

Vol 83 (5) ◽

pp. 783-793 ◽

Cited By ~ 5

Author(s):

B. Senowbari-Daryan ◽

G. D. Stanley

Keyword(s):

North America ◽

Shallow Water ◽

Type Species ◽

Upper Triassic ◽

Northern Calcareous Alps ◽

Late Triassic ◽

Separate Species ◽

Northern Calcareous ◽

Siliceous Spicules ◽

Horizontal Layers

Stromatomorpha californica Smith is a massive, calcified, tropical to subtropical organism of the Late Triassic that produced small biostromes and contributed in building some reefs. It comes from the displaced terranes of Cordilleran North America (Eastern Klamath terrane, Alexander terrane, and Wrangellia). This shallow-water organism formed small laminar masses and sometimes patch reefs. It was first referred to the order Spongiomorphidae but was considered to be a coral. Other affinities that have been proposed include hydrozoan, stomatoporoid, sclerosponge, and chambered sponge. Part of the problem was diagenesis that resulted in dissolution of the siliceous spicules and/or replaced them with calcite. Well-preserved dendroclone spicules found during study of newly discovered specimens necessitate an assignment of Stromatomorpha californica to the demosponge order Orchocladina Rauff. Restudy of examples from the Northern Calcareous Alps extends the distribution of this species to the Tethys, where it was an important secondary framework builder in Upper Triassic (Norian-Rhaetian) reef complexes. Revisions of Stromatomorpha californica produce much wider pantropical distribution, mirroring paleogeographic patterns revealed for other tropical Triassic taxa. Review of Liassic material from the Jurassic of Morocco, previously assigned to Stromatomorpha californica Smith var. columnaris Le Maitre, cannot be sustained. Species previously included in Stromatomorpha are: S. stylifera Frech (type species, Rhaetian), S. actinostromoides Boiko (Norian), S. californica Smith (Norian), S. concescui Balters (Ladinian-Carnian), S. pamirica Boiko (Norian), S. rhaetica Kühn (Rhaetian), S. stromatoporoides Frech, and S. tenuiramosa Boiko (Norian). Stromatomorpha rhaetica Kühn described from the Rhaetian of Vorarlberg, Austria shows no major difference from S. californica. An example described as S. oncescui Balters from the Ladinian-Carnian of the Rarau Mountains, Romania, is very similar to S. californica in exhibiting similar spicule types. However, because of the greater distance between individual pillars, horizontal layers, and the older age, S. oncescui is retained as a separate species. The net-like and regular skeleton of Spongiomorpha sanpozanensis Yabe and Sugiyama, from the Upper Triassic of Sambosan (Tosa, Japan), suggests a closer alliance with Stromatomorpha, and this taxon possibly could be the same as S. californica.

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A new riparian Mantidactylus (Brygoomantis) frog from the Tsaratanana and Manongarivo Massifs in northern Madagascar

Zootaxa ◽

10.11646/zootaxa.4486.4.10 ◽

2018 ◽

Vol 4486 (4) ◽

pp. 575 ◽

Cited By ~ 4

Author(s):

MIGUEL VENCES ◽

ANDREA HILDENBRAND ◽

KATHARINA M. WARMUTH ◽

FRANCO ANDREONE ◽

FRANK GLAW

Keyword(s):

New Species ◽

Nuclear Dna ◽

Taxonomic Revision ◽

Morphological Characters ◽

Molecular Data ◽

Test Case ◽

Nominal Species ◽

Type Specimens ◽

The Status ◽

Morphological Differences

The subgenus Brygoomantis in the Madagascar-endemic genus Mantidactylus contains 12 nominal species but is in urgent need of taxonomic revision as many additional, genetically divergent but undescribed candidate species have been identified. We here take a first step towards a better resolution of this group by describing a new species, Mantidactylus schulzi sp. nov., occurring at the Tsaratanana and Manongarivo Massifs, differentiated in genetic, bioacoustic and sometimes morphological characters from its closest relatives. We show that upon detailed study, most species in Brygoomantis can be delimited by concordant differentiation of mitochondrial and nuclear DNA, and by bioacoustic and morphological differences. We flag this group of morphologically similar frogs as a test case where molecular data on historical type specimens by ancient DNA methods might be needed to reach a satisfying clarification of taxonomy and nomenclature. However, the status of the new species M. schulzi is not in doubt as it is morphologically distinct from most historical type specimens, and microendemic to a region in northern Madagascar from where no earlier names exist.

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Systematic revision of Calligrapha s. str. Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae)

Zootaxa ◽

10.11646/zootaxa.4953.1.1 ◽

2021 ◽

Vol 4953 (1) ◽

pp. 1-111

Author(s):

JESÚS GÓMEZ-ZURITA

Keyword(s):

Leaf Beetle ◽

South American ◽

The South ◽

South American Species ◽

Systematic Revision ◽

Generic Type ◽

The Status ◽

Taxonomic Changes ◽

Current Species

In this work, the Chrysomelinae leaf beetle subgenus Calligrapha s. str. Chevrolat, 1836 is revised, offering redescriptions and keys for identification of twelve species currently considered in this group, allied to the South American species Calligrapha polyspila (Germar, 1821), the generic type of Calligrapha. The current species count results from important taxonomic changes. These include reversing a long-held synonymy, resurrecting the name Calligrapha mexicana Stål, 1859 stat. rev. for a species that is different from Chrysomela serpentina Rogers, 1856; upgrading the status of Polyspila serpentina var. discrepans Achard, 1923 to Calligrapha discrepans (Achard) stat. rev.; and formally proposing a number of new synonymies for several species, including: (1) Calligrapha discrepans (Achard) (= Calligrapha serpentina ssp. temaxensis Bechyné, 1952 syn. nov.); (2) Calligrapha fulvipes (Gistel, 1848) (= Calligrapha bajula Stål, 1860 syn. nov.; = Calligrapha nupta Stål, 1859 syn. nov.; = C. sponsa Stål, 1859 syn. nov.); and (3) Calligrapha polyspila (Germar) (= Polyspila polyspila var. bilineolata Achard, 1923 syn. nov.; = Polyspila polyspila var. plagata Achard, 1923 syn. nov.).

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Mesodiphthera Tillyard, 1919, from the Late Triassic of Queensland, the oldest cicada (Hemiptera: Cicadomorpha: Cicadoidea: Tettigarctidae)

Zootaxa ◽

10.11646/zootaxa.4567.2.8 ◽

2019 ◽

Vol 4567 (2) ◽

pp. 358 ◽

Cited By ~ 1

Author(s):

KEVIN J. LAMBKIN

Keyword(s):

Basal Cell ◽

Structural Diversity ◽

Wide Distribution ◽

Nodal Line ◽

Late Triassic ◽

Type Specimens ◽

Fossil Insect ◽

The Family ◽

New Material ◽

Cross Vein

New specimens of its type species from the Queensland Late Triassic (Norian) (~227–~208.5 Ma) fossil insect locality at Dinmore have revealed that the old and obscure Late Triassic genus Mesodiphthera Tillyard, 1919, from nearby Denmark Hill, is a tettigarctid cicada, the earliest record of the family and the oldest cicada. The genus is distinguished by the combined presence of three characters: the primary forks of R and M at about the same level, midway between the basal cell and the nodal line; RA2 with four or five terminal branches; and the inter-medial cross-vein backwardly inclined, running between M2 and M3. Of the three species originally ascribed to Mesodiphthera by Tillyard, only its type, M. grandis Tillyard, 1919, is retained in the genus. The other two species differ significantly from the type and are transferred to Tardilly gen. nov., which is similar to Mesodiphthera in the more or less aligned primary forks of R and M placed at about midway between the basal cell and the nodal line, and the backwardly inclined inter-medial cross-vein which runs between M2 and M3. It differs, however, in its smaller size, broader costal space, three-branched M3+4, and differently shaped CuA and CuA2. The new material, all of which is of M. grandis, provides a complete picture of the shape, colour and venation of its tegmen, whereas Tardilly prosboloides (Tillyard) comb. nov., 1922 and Tardilly dunstani (Tillyard) comb. nov., 1922 are still known only from their poorly preserved type specimens. Mesodiphthera and Tardilly exhibit a number of presumed plesiomorphies, viz the costal space much wider than the CuA cell, the basal cell strongly narrowed apically, and the post-nodal cross-vein series closer to the nodal line than the apex, which place it in the probable paraphyletic subfamily Cicadoprosbolinae. A more informed assessment of their relationships, however, must await a comprehensive analysis of the now 29 fossil genera of the family. The Tettigarctidae were the only cicadas of the Mesozoic and the discovery in the Triassic of Australia of Mesodiphthera and Tardilly clearly distinct from the 24 previously known Mesozoic genera, further demonstrates the family’s high degree of structural diversity, and emphasises its almost world-wide distribution in that Era.

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Morphological and molecular characters reveal the status of Pedesta rubella (Devyatkin, 1996) stat. n. and P. similissima (Devyatkin, 2002) syn. n. (Lepidoptera, Hesperiidae)

Zootaxa ◽

10.11646/zootaxa.4743.2.5 ◽

2020 ◽

Vol 4743 (2) ◽

pp. 217-231

Author(s):

MENG LI ◽

ALEXANDER L. MONASTYRKII ◽

KIRILL A. KOLESNICHENKO ◽

ZIHAO LIU ◽

GUOXI XUE ◽

...

Keyword(s):

Neighbor Joining ◽

Distribution Map ◽

Separate Species ◽

Outgroup Species ◽

Wing Patterns ◽

The Status ◽

Nj Tree ◽

The Mean ◽

Coi Sequences ◽

Molecular Characters

To clarify the relationships of Pedesta submacula (Leech, 1890), P. submacula rubella (Devyatkin, 1996) and P. similissima (Devyatkin, 2002), specimens from various localities in China and Vietnam, including the type materials of P. submacula rubella and P. similissima were examined. A neighbor-joining (NJ) tree was also reconstructed based upon partial COI sequences of 24 ingroup specimens and 4 outgroup species. The result shows that P. similissima is a synonym of P. submacula, and P. submacula rubella belongs to a separate species. The mean Kimura-2-Parameter genetic distance between P. submacula and P. rubella stat. n. is 4.4%. Wing patterns and genitalic structures of these taxa are illustrated and compared. A distribution map of both species is provided.

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