Locomotion of the Limpet, Patella Vulgata L

1970 ◽  
Vol 52 (1) ◽  
pp. 201-216
Author(s):  
H. D. JONES ◽  
E. R. TRUEMAN

1. The locomotion of Patella has been studied by histological, photographic and experimental techniques. The foot consists principally of dorso-ventral and transverse muscles and has no longitudinal muscle fibres near to its sole. The pedal haemocoel is limited to a region of small spherical cavities, extending for 0.5 mm. above the sole, two pedal sinuses and several lateral vertical channels. 2. Patella moves forward by means of retrograde alternate ditaxic pedal locomotory waves, but during rarely observed backward movement the waves are direct. During tight turns the limpet uses forward locomotion on one half of the foot and backward locomotion on the other. 3. During the passage of a retrograde locomotory wave the foot is lifted off the substrate by about 0.2 mm. and the pressure beneath the sole falls by about 6 cm. of water. 4. A model is proposed to account for locomotion utilizing the dorso-ventral muscles as the sole propulsive agents in the hydraulic system of the foot. This system consists principally of the dorso-ventral and transverse muscles, the spherical cavities of the pedal haemocoel and the fluid enclosed beneath each locomotory wave. Both fluid systems may be utilized during contraction and relaxation of different groups of dorso-ventral muscles.

1957 ◽  
Vol 3 (1) ◽  
pp. 111-122 ◽  
Author(s):  
Jean Hanson

1. The structure of the smooth muscle fibres in the longitudinal muscle coat of the body wall of Lumbricus terrestris has been investigated by phase contrast light microscopy and electron microscopy. 2. The muscle fibre is ribbon-shaped, and attached to each of its two surfaces is a set of myofibrils. These are also ribbon-shaped, and they lie with their surfaces perpendicular to the surfaces of the fibre, and their inner edges nearly meeting in the middle of the fibre. These fibrils are oriented at an angle to the fibre axis, and diminish greatly in width as they approach the edge of the fibre. The orientation of the set of fibrils belonging to one surface of the fibre is the mirror image of that of the set belonging to the other surface; thus, when both sets are in view in a fibre lying flat on one face, the fibre exhibits double oblique striation. A comparison of extended and contracted fibres indicates that as the fibre contracts, the angle made between fibre and fibril axes increases (e.g. from 5 to 30°) and so does the angle made between the two sets of fibrils (e.g. from 10 to 60°). 3. The myofibril, throughout its length, contains irregularly packed filaments, commonly 250 A in diameter, which are parallel to its long axis and remain straight in contracted muscles. Between them is material which probably consists of much finer filaments. Thus A and I bands are absent. 4. Bound to one face of each fibril, but not penetrating inside it, is a regularly spaced series of transverse stripes. They are of two kinds, alternating along the length of the fibril, and it is suggested that they are comparable to the Z and M lines of a cross-striated fibril. The spacing of these stripes is about 0.5 µ ("Z" to "Z") in extended muscles, and 0.25 µ in contracted muscles. A bridge extends from each stripe across to the stripeless surface of the next fibril.


1980 ◽  
Vol 87 (1) ◽  
pp. 121-136 ◽  
Author(s):  
J. H. Koenig ◽  
K. Ikeda

Simultaneous intracellular recordings were made from the six ipsilateral dorsal longitudinal muscle fibres of Drosophila in stationary flight. The influence of the firing of one motor unit upon the firing of another was analysed by observing the relationship between the interspike interval of a unit and the relative firing times of the other motor units within that interval. The analysis suggests that the influence is insignificant except when one unit would have fired soon after another. Then, a neural interaction occurs that can cause a unit to fire either earlier or later, depending on its firing relationship with the other units. Thus, the observation that no DLM fibre fires soon after another is the result of both a delaying effect and an effect which causes a cell to fire earlier than it normally would have fired.


Author(s):  
Shi Yan ◽  
Binbin He ◽  
Naizhi Zhao

Pipeline structure may generate damages during its service life due to the influence of environment or accidental loading. The damages need to be detected and repaired if they are severe enough to influence the transportation work. Non-destructive detection using smart materials combined with suitable diagonal algorithms are widely used in the field of structural health monitoring (SHM). Piezoelectric ceramics (such as Lead Zirconate Titanate, PZT) is one of the smart materials to be applied in the SHM due to the piezoelectric effect. So far, the PZT-based wave method is widely used for damage detection of structures, in particular, pipeline structures. A series of piezoelectric patches are bonded on the surface of the pipeline structure to monitor the damages such as local crack or effective area reduction due to corrosion by using diagonal waves. The damage of the pipeline structure can be detected by analysis of the received diagonal waves which peak value, phase, and arriving time can be deferent from the health ones. The response of the diagonal wave is not only correlated to the damage location through estimation of the arrival time of the wave peak, but also associated with the peak value of the wave for the reduction of wave energy as the guided wave passing through the damages. Therefore, the presence of damages in the pipeline structure can be detected by investigating the parameter change of the guided waves. The change of the wave parameters represents the attenuation, deflection and mode conversion of the waves due to the damages. In addition, the guided wave has the ability of quick detecting the damage of the pipeline structure and the simplicity of generating and receiving detection waves by using PZT patches. To verify the proposed method, an experiment is designed and tested by using a steel pipe bonded the PZT patches on the surface of it. The PZT patches consist of an array to estimate the location and level of the damage which is simulated by an artificial notch on the surface of the structure. The several locations and deep heights of the notches are considered during the test. A pair of the PZT patches are used at the same time as one is used as an actuator and the other as a sensor, respectively. A tone burst of 5 cycles of wave shape is used during the experiment. A wave generator is applied to create the proposed waves, and the waves are amplified by an amplifier to actuate the PZT patch to emit the diagonal waves with appropriately enough energy. Meanwhile, the other PZT patch is used as a sensor to receive the diagonal signals which contain the information of the damages for processing. For data processing, an index of root mean square deviation (RMSD) of the received data is used to estimate the damage level by compare of the data between the damaged and the health peak valves of the received signals. The time reversal method which aimed at increasing the efficiency of the detection is also used to detect the damage location by estimating the arrival time of the reflected wave passing with a certain velocity. The proposed method experimentally validates that it is effective for application in damage detection of pipeline structure.


It was shown in an earlier paper (7) that if maximal stimulation of either of two different afferent nerves can reflexly excite fractions of a given flexor muscle, there are generally, within the aggregate of neurones which innervate that muscle, motoneurones which can be caused to discharge by either afferent (i. e., motoneurones common to both fractions). The relationship which two such afferents bear to a common motoneurone was shown, by the isometric method of recording contraction, to be such that the activation of one afferent, at a speed sufficient to cause a maximal motor tetanus when trans­mitted to the muscle fibres, caused exclusion of any added mechanical effect when the other afferent was excited concurrently. This default in mechanical effect was called “occlusion.” Occlusion may conceivably be due to total exclusion of the effect of one afferent pathway on the common motoneurone by the activity of the other; but facilitation of the effect of one path by the activation of the other when the stimuli were minimal suggests that, in some circumstances at least, the effect of each could augment and summate with th at of the other at the place of convergence of two afferent pathways. Further investigation, using the action currents of the muscle as indication of the nerve impulses discharged by the motoneurone units, has now given some information regarding the effect of impulses arriving at the locus of convergence by one afferent path when the unit common to both is already discharging in response to impulses arriving by the other afferent path. Our method has been to excite both afferent nerves in overlapping sequence by series of break shocks at a rapid rate and to examine the action currents of the resulting reflex for evidence of the appearance of the rhythm of the second series in the discharge caused by the first when the two series are both reaching the motoneurone.


Babel ◽  
1994 ◽  
Vol 40 (1) ◽  
pp. 21-37
Author(s):  
Ivo R.V. Hoefkens

Marguerite Yourcenar, known as an author, is also the translator of about a dozen works. My purpose here is to trace the evolution of her oeuvre in the field of translation in relation to her literary output. I have divided the former into three distinct periods, the first of which covers the closing years of the 1930s, when Marguerite Yourcenar translated Virginia Woolf's The Waves and Henry James's What Maisie Knew. Her interest in these authors is to a large extent stylistic. On the other hand, the translation of Constantin Cavafy's poetry, which was begun during the same period, reflects the intimist themes to be found in Marguerite Yourcenar's early narratives {Alexis and the others), although she was then already seeking out other thematic sources. The translation was only published in 1958. It consequently falls within a second period: that of the "présentations critiques" (critical commentaries). These major efforts in translation {Présentation critique de Constantin Cavafy, La Couronne et la Lyre, Fleuve profonde, Sombre rivière) are marked by a manifest preoccupation with the aesthetic. But themes of a more universal character and engagement in the socio-political sphere also enter into the choice of the texts for translation (negro spirituals, Présentation critique d'Hortense Flexner). These translations were contemporaneous with the creation of Marguerite Yourcenar's most important novels, namely Mémoires d'Hadrien and L'OEuvre au Noir. The last of the three periods, the 1980s, finds her tackling far less ambitious projects, the function of which tends increasingly towards ethical communication. The only one of them that bears any resemblance to the "présentations critiques" is the essay on Yukio Mishima and the translation of Cinq Nô Modernes, assuming that these are to be considered as an ensemble. Here, as elsewhere, it also emerges that Marguerite Yourcenar is largely indifferent to the existence of other translations.


1940 ◽  
Vol 30 (2) ◽  
pp. 139-178
Author(s):  
J. Emilio Ramirez

Summary Over a period of six months, from July to December, 1938, an investigation on microseismic waves has been carried out in the Department of Geophysics of St. Louis University. Four electromagnetic seismographs, specially designed for recording microseisms, were installed in the city of St. Louis in the form of a triangular network. Two of these were E-W components, one at the St. Louis University Gymnasium and the other 6.4 km. due west at Washington University. The other two were arranged as N-S components, one at the St. Louis University Gymnasium and one 6.3 km. due south at Maryville College. The speed of the photographic paper was 60 mm/min., and time signals were recorded automatically and simultaneously on each paper from the same clock every minute and at shorter intervals from a special pendulum and “tickler” combination by means of telephone wires. The results have demonstrated beyond doubt that microseismic waves are traveling and not stationary waves. The same waves have been identified at each one of the stations of the network, and also at Florissant, 21.8 km. away from St. Louis University. The speed of microseismic waves at St. Louis was determined from several storms of microseisms and it was found to be 2.67±0.03 km/sec. The direction of microseisms was also established for most of the storms and it was found that about 80 per cent of incoming microseisms at St. Louis were from the northeast quadrant during the interval from July to December, 1938. No microseisms were recorded from the south, west, or southwest. The period of the waves varied between 3.5 and 7.5 sec. The average period was about 5.4 sec. The microseismic wave length was therefore of the order of 14¼ km. A study of the nature of microseismic waves from the three Galitzin-Wilip components of the Florissant station reveals in the waves many of the characteristics of the Rayleigh waves; that is, the particles in the passage of microseismic waves move in elliptical orbits of somewhat larger vertical axis and with retrograde motion. A comparison carried over a period of more than a year between microseisms and microbarometric oscillations recorded by specially designed microbarographs showed no direct relationship between the two phenomena in wave form, group form, period, or duration of storms. The source of microseisms is to be found not over the land, but rather out over the surface of the ocean. The amplitudes of microseisms depend only on the intensity and widespread character of barometric lows traveling over the ocean. Several correlations between the two phenomena seem to make this conclusion rather evident. Special emphasis is laid on the fact that all the determined directions of incoming microseisms at St. Louis point to a deep barometric low over the ocean. The period of microseisms seems to be a function of the distance between the station and the source of microseisms. The exact mechanism by which barometric lows over the ocean water result in the production of microseisms needs further investigation. Large microseisms have been produced without any indication of surf near the coasts, or with winds blowing from the land toward the ocean.


2014 ◽  
Vol 758 ◽  
Author(s):  
S. Karimpour Ghannadi ◽  
Vincent H. Chu

AbstractNumerical simulations of the transverse dam-break waves (TDWs) produced by the sudden removal of a gate on the side of a waterway are conducted based on the shallow-water equations to find solutions to a family of water-diversion problems. The Froude numbers in the main flow identify the members of the family. The depth and discharge profiles are analysed in terms of Ritter’s similarity variable. For subcritical main flow, the waves are comprised of a supercritical flow expansion followed by a subcritical outflow. For supercritical main flow, on the other hand, the waves are analogous to the Prandtl–Meyer expansion in gas dynamics. The diversion flow rate of two-dimensional TDWs on a flat bed is 55 % greater than the one-dimensional flow rate of Ritter in the limiting case of zero main flow, and approaches the rate of Ritter in the other limit when the value of the Froude number in the main flow approaches infinity. The diversion flow rate over a weir is generally higher than the rate over a flat bed depending on the Froude number of the main flow. These numerical simulation results are consistent with laboratory observations.


1835 ◽  
Vol 125 ◽  
pp. 355-358 ◽  

The Fourth Memoir, published in my Zoological Researches and Illustrations, No. III. page 69, &c., having first made known the real nature of the Cirripedes , the key of which remained concealed in their metamorphosis, it might have been expected that some naturalist favourably situated to investigate the oceanic tribe of these animals, would have been the first to make the same discovery in regard to these, and thereby complete their natural history. It was scarcely to be expected that the honour of this discovery also should be reserved for the author, fixed to one spot, where none of them naturally exist, and are but casually thrown upon our shores by the waves of the Atlantic, attached to pieces of wreck, or brought into port fixed to the bottoms of ships returning from distant voyages. Fortunately, however, two ships of this description came into this harbour (Cork), one from the Mediterranean, the other from North America, which, not being sheathed with copper, had their bot­toms literally covered with Barnacles of the three genera of Lepas , Cineras , and Otion ; and having persons employed expressly for the purpose, numbers of these were brought alive in sea water, amongst which were many with the ova in various stages of their progress, and some ready to hatch, which they eventually did in prodigious numbers, so as to enable him to add the proof of their being, like the Balani, natatory Crusta­cea in their first stage , but of a totally different facies and structure; a circumstance which determines the propriety of the separation of the Cirripedes into two tribes, and evinces the sagacity of Mr. MacLeay in being the first to indicate that these two tribes, the Balani and Lepades , were not so closely related as generally supposed. The larvæ of the Balani , described in Memoir IV. under the external appearance of the bivalve Monoculi ( Astracoda ), have a pair of pedunculated eyes, more numerous and more completely developed members, approximating to those of Cyclops , and of the perfect Triton ; while, in the present type, or Lepades , the larva resembles some­what that of the Cyclops , which Müller, mistaking for a perfect animal, named Amymone , and which can be shown to he common to a great many of the Entomostraca ; or the resemblance is still more striking to that of the Argulus Armiger of Latreille, which, in fact, is but an Amymone furnished with a tricuspidate shield at the back.


1968 ◽  
Vol 3 (1) ◽  
pp. 49-54
Author(s):  
R. MILEDI ◽  
c. R. SLATER

An electron-microscopic study was made of the mitochondria in normal and denervated rat diaphragm muscles. Two types of structurally different muscle fibres were found: one type had a well-defined M-line, while the other did not. The mitochondria in normally innervated muscle are regularly arranged at both sides of the Z-line. The mitochondria are very long and branched, and surround the myofibrils at the level of the Z-line. In transverse sections of the muscle fibres the worm-like mitochondria give a very characteristic picture. After denervation the mitochondria are smaller and less regularly arranged. In transverse sections of the muscle fibres the mitochondria have small circular profiles. This contrasts sharply with the normal appearance and makes it possible to distinguish normal from denervated fibres. The mitochondrial changes can be detected less than 24 h after denervation.


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