scholarly journals Primed Acclimation of Papaya Increases Short-term Water Use But Does Not Confer Long-term Drought Tolerance

HortScience ◽  
2017 ◽  
Vol 52 (3) ◽  
pp. 441-449 ◽  
Author(s):  
Christopher Vincent ◽  
Diane Rowland ◽  
Bruce Schaffer
Keyword(s):  
Drought Stress ◽  
Soil Water ◽  
Water Deficit ◽  
Water Use ◽  
Dry Matter ◽  
Leaf Gas Exchange ◽  
Severe Drought ◽  
Dry Matter Accumulation ◽  
Short Term ◽  
Soil Water Tension

Primed acclimation (PA) is a regulated deficit irrigation (RDI) strategy designed to improve or maintain yield under subsequent drought stress. A previous study showed photosynthetic increases in papaya in response to a PA treatment. The present study was undertaken to test the duration of the PA effect when papaya plants were challenged with severe drought stress. Potted plants were stressed at 1, 2, and 3 months after conclusion of a PA treatment consisting of 3 weeks at soil water tension (SWT) of −20 kPa. Measurements included leaf gas exchange, root growth, and organ dry mass partitioning. PA did not reduce net CO2 assimilation (A) during the deficit period. At the end of the PA period, total dry matter accumulation per plant and for each organ was unaffected, but proportional dry matter partitioning to roots was favored. After resuming full irrigation, A increased and whole plant water use was more than doubled in PA-treated plants. However, water use and A of PA-treated plants decreased to reconverge with those of control plants by 6 weeks after the PA treatment. Over the course of the study, PA plants maintained lower stem height to stem diameter ratios, and shorter internode lengths. However, these changes did not improve photosynthetic response to any of the water-deficit treatments. We conclude that papaya exhibits some signs of stress memory, but that rapid short-term acclimation responses dominate papaya responses to soil water deficit.

Water use of wheat, barley, canola, and lucerne in the high rainfall zone of south-western Australia

2005 ◽  
Vol 56 (7) ◽  
pp. 743 ◽  
Author(s):  
Heping Zhang ◽  
Neil C. Turner ◽  
Michael L. Poole
Keyword(s):  
Soil Water ◽  
Water Use ◽  
Western Australia ◽  
Dry Matter ◽  
Grain Filling ◽  
Rooting Depth ◽  
Dry Matter Accumulation ◽  
High Rainfall ◽  
Annual Crops ◽  
Significant Difference

Water use of wheat (Triticum aestivum L.), barley (Hordeum vulgare L.), canola (Brassica napus L.), and lucerne (Medicago sativa L.) was measured on a duplex soil in the high rainfall zone (HRZ) of south-western Australia from 2001 to 2003. Rainfall exceeded evapotranspiration in all years, resulting in transient perched watertables, subsurface waterlogging in 2002 and 2003, and loss of water by deep drainage and lateral flow in all years. There was no significant difference in water use among wheat, barley, and canola. Lucerne used water at a similar rate to annual crops during the winter and spring, but continued to extract 80−100 mm more water than the annual crops over the summer and autumn fallow period. This resulted in about 50 mm less drainage past the root-zone than for annual crops in the second and third years after the establishment of the lucerne. Crop water use was fully met by rainfall from sowing to anthesis and a significant amount of water (120−220 mm) was used during the post-anthesis period, resulting in a ratio of pre- to post-anthesis water use (ETa : ETpa) of 1 : 1 to 2 : 1. These ratios were lower than the indicative value of 2 : 1 for limited water supply for grain filling. High water use during the post-anthesis period was attributed to high available soil water at anthesis, a large rooting depth (≥1.4 m), a high proportion (15%) of roots in the clay subsoil, and regular rainfall during grain filling. The pattern of seasonal water use by crops suggested that high dry matter at anthesis did not prematurely exhaust soil water for grain filling and that it is unlikely to affect dry matter accumulation during grain filling and final grain yield under these conditions.

scholarly journals Effect of Glyphosate on Guineagrass Submitted to Different Soil Water Potential

2018 ◽  
Vol 36 (0) ◽  
Author(s):  
A.C. SILVA JR. ◽  
C.G. GONÇALVES ◽  
M.C. SCARANO ◽  
M.R.R. PEREIRA ◽  
D. MARTINS
Keyword(s):  
Water Management ◽  
Soil Water ◽  
Water Deficit ◽  
Dry Matter ◽  
Soil Water Potential ◽  
Physiological Parameters ◽  
Dry Matter Accumulation ◽  
Intermediate Water ◽  
Water Restriction ◽  
Environment Temperature

ABSTRACT: The action of herbicides on weeds that develop under water deficit can be compromised, because the routes of penetration of hydrophilic herbicides are reduced due to the lower hydration of the cuticle in these plants. Moreover, hydrophobic compounds found in the epicuticular wax coating of plants under water stress, hinder the penetration of hydrophilic compounds. This study evaluated the control efficiency of glyphosate on guineagrass plants when submitted to different water deficits. The study was conducted in a greenhouse, and treatments were composed of guineagrass plants submitted to three soil water conditions [low water deficit (13%), intermediate water deficit (10%) and high water restriction (8%)], three doses of glyphosate (0.0, 270.0 and 540.0 g ha-1) and two phenological stages of plant development (4-6 leaves and 1-3 tillers). The water management started when plants presented two developed leaves. Visual evaluations were performed 7, 14, 21 and 35 days after the application of the herbicide; the morpho-physiological parameters of the specific leaf area, stomatal conductance and the difference between the environment temperature and the leaf temperature on the day of the herbicide application were also analyzed at the end of the study, as well as the dry matter of shoot and root. With the increase of the water restriction, there was a decrease in the analyzed morpho-physiological parameters, as well as in the dry matter accumulation of the shoot and roots of the studied plants. Plant control was more efficient when a 540 g ha-1 dose of glyphosate was applied, and when they were controlled at their vegetative stage of 1-3 tiller, and with a water management of 13%. It is possible to state that guineagrass under water restriction have less control efficacy when treated with glyphosate.

Tropical rainforests under severe drought stress: distinct water use strategies among and within species

2021 ◽  
Author(s):  
Angelika Kübert ◽  
Kathrin Kühnhammer ◽  
Ines Bamberger ◽  
Erik Daber ◽  
Jason De Leeuw ◽  
...  
Keyword(s):  
Gas Exchange ◽  
Soil Water ◽  
Water Use ◽  
Leaf Gas Exchange ◽  
Physiological Responses ◽  
Leaf Water ◽  
Severe Drought ◽  
Subsequent Recovery ◽  
Species Specific

<p>Increasing drought in the tropics is a major threat to rainforests and can strongly harm plant communities. Understanding species-specific water use strategies to drought and the subsequent recovery is therefore important for estimating the risk to tropical rainforest ecosystems of drought. Conducting a large-scale long-term drought experiment in a model rainforest ecosystem (Biosphere 2 WALD project), we evaluated the role of plant physiological responses, above and below ground, in response to drought and subsequent recovery in five species (3 canopy species, 2 understory species). The model rainforest was exposed to a 9.5-week lasting drought. Severe drought was ended with a deep water pulse strongly enriched in <sup>2</sup>H, which allowed us to distinguish between deep and shallow rooting plants, and subsequent rain (natural abundance range of <sup>2</sup>H). We assessed plant physiological responses by leaf water potential, sap flow and high resolution monitoring of leaf gas exchange (concentrations and stable isotopes of H<sub>2</sub>O and CO<sub>2</sub>). Thereby, we could derive plant water uptake and leaf water use efficiency (WUE<sub>leaf</sub>) in high temporal resolution, revealing short-term and long-term responses of plant individuals to drought and rewetting. The observed water use strategies of species and plants differed widely. No uniform response in assimilation (A) and transpiration (T) to drought was found for species, resulting in decreasing, relatively constant, or increasing WUE<sub>leaf</sub> across plant individuals. While WUE<sub>leaf</sub> of some plant individuals strongly decreased due to a breakdown in A, others maintained relatively high T and A and thus constant WUE<sub>leaf, </sub>or increased WUE<sub>leaf</sub> by decreasing T while keeping A relatively high. We expect that the observed plant-specific responses in A, T and WUE<sub>leaf</sub> were strongly related to the plant individuals' access to soil water. We assume that plant individuals with constant WUE<sub>leaf</sub> could maintain their leaf gas exchange due to access to water of deeper soil layers, while plants with increasing/decreasing WUE<sub>leaf</sub> mainly depended on shallow soil water and only had limited or no access to deep soil water. We conclude that the observed physiological responses to drought were not only determined by species-specific water use strategies but also by the diverse strategies within species, mainly depending on the plant individuals' size and place of location. Our results highlight the plasticity of water use strategies beyond species-specific strategies and emphasize its importance for species’ survival in face of climate change and increasing drought.</p>

Adaptation to and recovery from drought stress at vegetative stages in wheat (Triticum aestivum) cultivars

2016 ◽  
Vol 43 (12) ◽  
pp. 1159 ◽  
Author(s):  
Muhammad Abid ◽  
Zhongwei Tian ◽  
Syed Tahir Ata-Ul-Karim ◽  
Feng Wang ◽  
Yang Liu ◽  
...  
Keyword(s):  
Water Stress ◽  
Drought Stress ◽  
Grain Yield ◽  
Dry Matter ◽  
Leaf Gas Exchange ◽  
Field Capacity ◽  
Severe Drought ◽  
Drought Intensity ◽  
Crop Growth Rate ◽  
Physiological Adaptations

Studying plants’ capability to adapt and recover from drought stress is essential because of the ever-changing nature of drought events. To evaluate the genotypically variable morpho-physiological adaptations to drought stress and recovery after re-watering, two wheat cultivars (Luhan-7 and Yangmai-16) were pot-cultured under three levels of water stress: severe (35–40% field capacity, FC) and moderate water deficits (55–60% FC) and well-watered conditions. Drought stress was applied at tillering (Feekes 2 stage) and jointing (Feekes 6 stage), respectively, followed by re-watering, and we observed changes in leaf characteristics, growth and physiological activities during water stress and rewatering periods as well as final grain yield traits at maturity. Results showed that drought stress adaptability associated with reduced leaf area, higher leaf thickness, chlorophyll, leaf dry matter and maintenance of leaf water potential were more strongly pronounced in Luhan-7 than in Yangmai-16. Under moderate stress both cultivars exhibited a small decrease in leaf gas-exchange and chlorophyll fluorescence activities, followed by rapid recovery. Under severe drought stress, Yangmai-16 displayed relatively less adaptability to drought, with a slower recovery after re-watering and a greater decrease in grain yield. It was concluded that even though crop growth rate completely recovered after re-watering, the final dry matter and grain yield outcomes were affected by pre-drought stress, and were dependant on the drought intensity, adaptability and recovery differences of the cultivars and growth stage. It was also concluded that genotypic variations in adaptability and recovery from drought stress are the indicators of drought tolerance and grain yield sustainability in wheat.

scholarly journals IRRIGATION AND MULCHING MANAGEMENT FOR SWEET PEPPER CROP IN PROTECTED ENVIRONMENT

Irriga ◽  
2004 ◽  
Vol 9 (3) ◽  
pp. 217-224 ◽  
Author(s):  
Antonio Evaldo Klar ◽  
Sidnei Osmar Jadoski ◽  
Sidnei Osmar Jadoski
Keyword(s):  
Drought Stress ◽  
Soil Water ◽  
Capsicum Annuum ◽  
Water Use ◽  
Sweet Pepper ◽  
Fruit Yield ◽  
Severe Drought ◽  
Capsicum Annuum L ◽  
Polyethylene Mulching ◽  
Engineering Department

IRRIGATION AND MULCHING MANAGEMENT FOR SWEET PEPPER CROP IN PROTECTED ENVIRONMENT   Antonio Evaldo Klar1; Sidnei Osmar Jadoski21Rural Engineering Department, Agronomic Science College,Paulista State University ,  Botucatu-SP (Brazil),  [email protected] and Veterinarian Sciences Center,  Guarapuava – PR.    1 ABSTRACT  This study was developed over the 1999-2000 agricultural years at the Rural Engineering Department of School of Agronomical Sciences, UNESP, Botucatu, SP, under protected environment. The objective was to evaluate the influence of irrigation and black polyethylene mulch management on the water use and fruit production of sweet pepper crop (Capsicum annuum L., Elisa Hybrid) 230 days after seedling transplant (DAST). The study was divided into two experiments: 1) application of the irrigation at -50 and -1500 kPa minimum soil water potentials with and without mulching on the soil, from 29 to 168 DAST. A randomized experimental design was used with six replications; and 2) severe water deficit in the soil through irrigation suspension and mulching removal (169 to 230 DAST). Drip fertigation was used and soil water was monitored by tensiometers and neutron probe. It was verified that: - the severe drought stress caused leaf senescence and abscission and significantly affected the production and quality of sweet pepper fruits and the Water Use Efficiency (WUE); - the fruit yield and number were higher in the treatments with mulching, therefore polyethylene mulching showed to be an efficient technique to reduce irrigation number and water volume applied. This efficiency was reduced with soil water content decreases based on fruit yield. Pepper plants showed good osmotic adjustment and, consequently, tolerance to water stress.   KEYWORDS: drought stress, polyethylene mulching, soil water potential   KLAR, A. E.; JADOSKI, S. O. IRRIGAÇÃO E COBERTURA MORTA EM PLANTAS DE PIMENTÃO EM AMBIENTE PROTEGIDO   2 RESUMO  O trabalho foi desenvolvido no ano agrícola 1999-2000, em casa de vegetação, no Departamento de Engenharia Rural da Faculdade de Ciências Agronômicas - UNESP, campus de Botucatu, SP. O objetivo foi estudar o efeito de diferentes manejos da água de irrigação e de cobertura de polietileno preto na superfície do solo sobre o consumo de água e produção de frutos da cultura do pimentão (Capsicum annuum L, híbrido Elisa), por um período de 230 dias após o transplante das mudas (DAT). O trabalho foi composto por dois experimentos: I) aplicação dos manejos da  irrigação a 50 kPa e 1500 kPa, com e sem a presença de cobertura de polietileno preto sobre a superfície do solo, para o período de 29 a 168 DAT. O delineamento experimental foi inteiramente casualizado com seis repetições, e II) aplicação de deficiência hídrica severa no solo, através da suspensão das irrigações e retirada da cobertura de polietileno do solo, no período de 169 a 230 DAT. Concluiu-se que a deficiência hídrica afeta a qualidade e a produção dos frutos do pimentão. A utilização de cobertura de polietileno sobre o solo é técnica eficiente para reduzir a necessidade de irrigações e o volume de água a ser aplicado aos cultivos e sua eficiência é reduzida na medida em que diminui o conteúdo de água do solo. Houve significativa influencia no ajuste osmótico pela aplicação dos estresses hídricos nos tratamentos em que o potencial mínimo de água no solo atingiu 1500 kPa.  UNITERMOS: deficiência hídrica, potencial de água do solo, polietileno preto

scholarly journals Soil Water Deficit and Fertilizer Placement Effects on Root Biomass Distribution, Soil Water Extraction, Water Use, Yield, and Yield Components of Soybean [Glycine max (L.) Merr.] Grown in 1-m Rooting Columns

2021 ◽  
Vol 12 ◽  
Author(s):  
Michael Gebretsadik Gebre ◽  
Hugh James Earl
Keyword(s):  
Drought Stress ◽  
Drought Tolerance ◽  
Soil Water ◽  
Water Use ◽  
Yield Components ◽  
Dry Matter ◽  
Soil Depth ◽  
Mild Drought ◽  
Watering Treatment ◽  
Pod Number

Typical small-pot culture systems are not ideal for controlled environment phenotyping for drought tolerance, especially for root-related traits. We grew soybean plants in a greenhouse in 1-m rooting columns filled with amended field soil to test the effects of drought stress on water use, root growth, shoot growth, and yield components. There were three watering treatments, beginning at first flower: watered daily to 100% of the maximum soil water holding capacity (control), 75% (mild drought stress), or 50% (drought stress). We also tested whether applying fertilizer throughout the 1-m soil depth instead of only in the top 30 cm would modify root distribution by depth in the soil profile and thereby affect responses to drought stress. Distributing the fertilizer over the entire 1-m soil depth altered the root biomass distribution and volumetric soil water content profile at first flower, but these effects did not persist to maturity and thus did not enhance drought tolerance. Compared to the control (100%) watering treatment, the 50% watering treatment significantly reduced seed yield by 40%, pod number by 42%, seeds per pod by 3%, shoot dry matter by 48%, root dry matter by 53%, and water use by 52%. Effects of the 75% watering treatment were intermittent between the 50 and 100%. The 50% treatment significantly increased root-to-shoot dry matter ratio by 23%, harvest index by 17%, and water-use efficiency by 7%. Seed size was not affected by either fertilizer or watering treatments. More than 65% of the total root dry matter was distributed in the upper 20 cm of the profile in all watering treatments. However, the two drought stress treatments, especially the mild drought stress, had a greater proportion of root dry matter located in the deeper soil layers. The overall coefficient of variation for seed yield was low at 5.3%, suggesting good repeatability of the treatments. Drought stress imposed in this culture system affected yield components similarly to what is observed in the field, with pod number being the component most strongly affected. This system should be useful for identifying variation among soybean lines for a wide variety of traits related to drought tolerance.

scholarly journals Physiological Acclimation of Taxodium Hybrid ‘Zhongshanshan 118’ Plants to Short-term Drought Stress and Recovery

HortScience ◽  
2016 ◽  
Vol 51 (9) ◽  
pp. 1159-1166
Author(s):  
Qin Shi ◽  
Yunlong Yin ◽  
Zhiquan Wang ◽  
Wencai Fan ◽  
Jianfeng Hua
Keyword(s):  
Drought Stress ◽  
Water Content ◽  
Soil Water ◽  
Water Deficit ◽  
Thermal Dissipation ◽  
Protective Mechanism ◽  
Short Term ◽  
Chlorophyll Contents ◽  
As Stress ◽  
Physiological Acclimation

The physiological acclimation of Taxodium hybrid ‘zhongshanshan 118’ (T.118) plants to a progressive drought stress and drought-stressed to recovery treatment (DS-R) was investigated in this study. Plants of control (C) treatment were watered daily throughout the experiment. Results indicated that water deficit reduced stomatal conductance (gS) to improve water use efficiency (WUE) and, as a consequence, net photosynthetic rate (Pn), transpiration rate (Tr), and intercellular CO2 concentration (Ci) were also decreased in DS-R T.118 plants compared with C plants. These reductions became more significant with decreasing soil water availability. Correlation analysis showed gS was positively correlated (P < 0.01) with the soil water content as well as leaf relative water content (RWC). There was a tendency to accumulate proline, malondialdehyde (MDA), antioxidases, and membrane electrolyte leakage as stress intensity increased. Moreover, drought stress induced significant (P < 0.05) decline in total chlorophyll contents (Chlt) and increase of nonphotochemical quenching (NPQ) on day 8 as a photo-protective mechanism. Cluster analysis distinguished the adaption of T.118 plants to water deficit in two ways. First, photosynthesis was related to thermal dissipation, and second antioxidation was related to morphology and osmosis. Furthermore, tested parameters showed a reversed tendency and restored equivalently to C levels after 9 days of rewatering. These findings suggest that T.118 plants demonstrated considerable tolerance to short-term drought stress and recovery due to a high degree of plasticity in physiological acclimation.

scholarly journals Influence of Spring and Fall Drought Stresses on Growth and Gas Exchange during Stress and Posttransplant of Container-grown Magnoli ×soulangiana `Jane'

2002 ◽  
Vol 127 (1) ◽  
pp. 38-44 ◽  
Author(s):  
R. Thomas Fernandez ◽  
Robert E. Schutzki ◽  
Kelly J. Prevete
Keyword(s):  
Drought Stress ◽  
Stomatal Conductance ◽  
Gas Exchange ◽  
Leaf Gas Exchange ◽  
Growth Index ◽  
Severe Drought ◽  
Severe Stress ◽  
Flower Number ◽  
Short Term ◽  
Whole Plant

Responses of Magnolia ×soulangiana (Soul.-Bod.) `Jane' (`Jane' saucer magnolia) to consecutive short term pretransplant drought stresses and recovery after transplanting were evaluated beginning October 1997 and June 1998. Plants were subjected to one (mild) or two (moderate) 3-day drought stress periods or a two 3-day and one 4-day (severe) drought stress period, each separated by two rewatering periods over 24 hours. One day after each stress period, plants were transplanted into the field and well watered to monitor recovery from stress. Plant response was determined by measuring whole-plant CO2 assimilation, leaf gas exchange (CO2 assimilation, transpiration, stomatal conductance) and canopy growth throughout stress and recovery periods. Whole-plant and leaf CO2 assimilation were lower for the stressed treatments for most of the measurements taken during stress in the fall and spring. After release from stress and transplanting, leaf CO2 assimilation returned to control levels for mild and moderate fall stresses within 2 to 3 d by the next measurement, while it was over 3 weeks until recovery from the severe stress. There was no difference in leaf gas exchange following release from stress and transplanting during the spring stress. More rapid defoliation occurred for the severe fall-stressed plants compared to the controls after release from stress in the fall. Flower number was reduced in spring for the fall-stressed plants. At termination of the experiment, the growth index was lower for severe fall-stressed plants but there were no differences for other fall stress treatments. There was no increase in growth for control or stressed plants for the spring experiment.

Dry matter accumulation and water use relationships in wheat crops

1979 ◽  
Vol 30 (5) ◽  
pp. 815 ◽  
Author(s):  
AD Doyle ◽  
RA Fischer
Keyword(s):  
Soil Water ◽  
Water Use ◽  
Dry Matter ◽  
Plant Density ◽  
Sowing Date ◽  
Crop Growth ◽  
Soil Evaporation ◽  
Total Solar Radiation ◽  
Dry Matter Accumulation ◽  
Area Index

In order to better understand crop growth (dry matter accumulation, DM), crop evapotranspiration (Et), and their interrelationships, we studied dryland crops of wheat (Triticum aestivum L. cv. Timgalen) sown at various dates and seeding densities in each of three years at Tamworth, N.S.W. Soil water stress was minimal before anthesis in each year, but in two years substantial stress arose before maturity. DM was increased consistently by increased plant density, and decreased at anthesis and maturity by later sowing. Crop growth rates determined over 2-week intervals around anthesis ranged from 3 to 20 g m-2 d-1, representing a range in efficiency of utilization of intercepted total solar radiation of 0.48 to 2.35%, variation which was adequately explained (R2 = 0.80) by ontogeny (days from anthesis) and Et / Ep ratio (Ep = class A pan evaporation). Et at anthesis, but not at maturity, was increased slightly by higher seedmg density; crop Et was not consistently affected by sowing date. Et / Ep over 2-week periods around anthesis was related to leaf area index, and to a lesser extent to available soil water and Ep (R2 = 0.58). For the period from the first sowing date in June or July until the middle of October, the relationship of total Et to DM production was linear and close each year, but the slope varied from 6.2 g m-2 mm-1 (cold dry year) to 14.0 g m-2 mm-1 (wet year). This variation could be attributed to annual variation in the soil evaporation component of Et, and in the ratio of DM to crop transpiration (= transpiration efficiency, TE). For 2-week periods around anthesis, TE ranged from 2.9 to 5.4 g m-2 mm-1 and was inversely related to Ep (R2 = 0.56). Provided soil evaporation can be allowed for, since it ranged from 18 to 41% of crop Et from sowing to maturity, it is argued that the crop transpirationtranspiration efficiency approach is particularly useful for analysing the growth and water use of dryland wheat.

Water use by winter wheat as affected by soil management

1984 ◽  
Vol 103 (1) ◽  
pp. 189-199 ◽  
Author(s):  
M. J. Goss ◽  
K. R. Howse ◽  
Judith M. Vaughan-Williams ◽  
M. A. Ward ◽  
W. Jenkins
Keyword(s):  
Winter Wheat ◽  
Soil Water ◽  
Water Deficit ◽  
Water Use ◽  
Management Practices ◽  
Soil Management ◽  
Maximum Depth ◽  
Water Extraction ◽  
Soil Water Deficit ◽  
Potential Yield

SummaryIn each of the years from September 1977 to July 1982 winter wheat was grown on one or more of three clay soil sites (clay content 35–55%) in Oxfordshire where the climate is close to the average for the area of England growing winter cereals.The effects on crop water use of different soil management practices, including ploughing, direct drilling and subsoil drainage, are compared. Cultivation treatment had little effect on the maximum depth of water extraction, which on average in these clay soils was 1·54 m below the soil surface. Maximum soil water deficit was also little affected by cultivation; the maximum recorded value was 186±7·6 mm. Subsoil drainage increased the maximum depth of water extraction by approximately 15 cm and the maximum soil water deficit by about 17 mm.Generally soil management had little effect on either total water use by the crop which was found to be close to the potential evaporation estimated by the method of Penman, or water use efficiency which for these crops was about 52 kg/ha par mm water used.Results are discussed in relation to limitations to potential yield.

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