Studies of cattle and sheep eating leaf and stem fractions of grasses. 3. The retention time in the rumen of large feed particles

1981 ◽  
Vol 32 (1) ◽  
pp. 123 ◽  
Author(s):  
DP Poppi ◽  
DJ Minson ◽  
JH Ternouth

Pangola grass (Digitavia decumbens) and Rhodes grass (Chlovis gayana) cut as 6 and 12 week regrowths were separated into leaf and stem fractions and fed ad libitum to four cattle and eight sheep fitted with ruminal fistulae to determine the importance of particle size in controlling the retention time of feed in the rumeno-reticulum (rumen). Particle size was determined by using a wet sieving technique, and based on the cumulative dry matter distribution on the sieves of faeces from cattle and sheep; all particles >1.18 mm were described as large particles. The proportion of large particles was measured in the chopped diet offered, the masticated diet, the ruminal contents and the faeces. Chopped leaf and stem fractions contained 0.85 and 0.86 g/g large particles respectively. Mastication by cattle reduced the proportion of large particles in leaf and stem to 0.58 and 0.76 (P < 0.01), and mastication by sheep to 0.56 and 0.67 (P < 0.01). The proportion of large particles in the rumen of sheep was 0.236 and 0.249 for leaf and stem respectively (P > 0.05) and in cattle 0.272 and 0.345 (P < 0.05). The faeces contained relatively few large feed particles (< 0.045 for cattle and <0.018 for sheep). Cattle eating leaf and stem produced faeces containing 0.027 and 0.040 (g/g) large particles (P <0.05), compared with only 0,008 and 0.013 (P < 0.05) in sheep. Most of the large particles entering the rumen in the masticated feed disappeared in the rumen by breakdown to smaller particles or by digestion. In cattle the proportions that disappeared were 0.978 and 0.976 (P > 0.05) for leaf and stem, and in sheep 0.993 and 0.991 (P > 0.05). Large particles in leaf and stem were retained in the rumen of the sheep for 11.0 and 11.7 h (P > 0.05). Cattle retained large particles in all diets for a longer time (P < 0.01); 16.1 h for leaf and 20,2 h for stem (P > 0.05). Differences in retention time of large particles in the rumen did not appear to be the only factor controlling the retention of dry matter in the rumen and voluntary intake. A model was developed to describe the flow of large and small particles through the rumen. Intake simulation studies indicated that the most important factor influencing dry matter retention time in the rumen was the retention time of small particles (> 1.18 mm). Changes in the rate of breakdown of large particles had a small effect on dry matter retention time.


1981 ◽  
Vol 32 (1) ◽  
pp. 99 ◽  
Author(s):  
DP Poppi ◽  
DJ Minson ◽  
JH Ternouth

Pangola grass (Digitaria decumbens) and Rhodes grass (Chloris gayana) were cut as 6 and 12 week regrowths, dried, chopped, and separated into leaf and stem fractions by using a gravity separator. Each of the eight diets was offered to four cattle and eight sheep fitted with a ruminal cannula to measure voluntary intake and digestibility of the dry matter (DM), organic matter (OM) and neutral detergent fibre (NDF). Possible deficiencies of protein and minerals were minimized in this study by feeding casein and a mineral supplement. The total DM, OM and NDF contents of the rumeno-reticulum (rumen) were measured by manually emptying the rumen and the values used to estimate retention times of DM, OM and NDF. Cattle consumed 35% more leaf than stem fraction. Sheep consumed 21 % more leaf. When the daily voluntary intake was expressed as g/kg0.9 the intake was similar for cattle and sheep. The higher intake of the leaf fraction was associated with the shorter time that it was retained in the rumen of both cattle and sheep. Leaf and stem fractions were digested to the same extent by cattle. There was also no difference in the digestibility of leaf and stem fed to sheep. The digestive efficiency of sheep was 0.033 lower than that of cattle for both DM and NDF. Cattle retained all diets for longer periods in the rumen than did sheep. This difference was associated with the higher digestive efficiency of the cattle. It was concluded that both cattle and sheep consumed more leaf than stem fractions of grasses, and that the higher intake of leaf was associated with the shorter time that it was retained in the rumen and not by differences in digestibility as such.



1981 ◽  
Vol 32 (2) ◽  
pp. 389 ◽  
Author(s):  
RE Hendricksen ◽  
DP Poppi ◽  
DJ Minson

Hay made from the tropical forage legume Lablab purpureus was chopped and separated into leaf and stem fractions by using a gravity separator. Each diet was fed to four cattle and six sheep fitted with rumen fistulae to measure the digestibilities of the dry matter (DM), organic matter (OM) and neutral detergent fibre (NDF), and the retention times of DM, OM and NDF in the rumeno-reticulum (rumen) and the rumen volume. Mean voluntary intake of the DM in the leaf fraction was 79 % higher by cattle and 61 % higher by sheep than the stem fraction, and this appeared to be associated with the shorter time it was retained in the rumen. Both leaf and stem DM were digested to a similar extent by cattle. Sheep digested the leaf fraction to the same extent as cattle. However, the DM digestibility of the stem eaten by sheep was 0,061 lower than that of the leaf fraction (P < 0.05). Cattle retained the DM, OM and NDF in all diets for longer periods in the rumen than did sheep. This difference led to a 48% higher voluntary intake (g(kg body d-1) by sheep. The proportion of large particles that disappeared within the rumen was lower (P < 0.01) for cattle (0.968) than for sheep (0.994). The large particles of the leaf and stem fraction were retained for the same mean time by sheep (5.7 and 5.6 h), which was shorter (P < 0.01) than the corresponding times for cattle (13.8 and 21.6 h). The rate of disappearance of large particles in the rumen appeared to be only a minor factor causing the longer retention time of stem fractions in the rumen. The concentrations of volatile fatty acids and ammonia in the rumen were higher for leaf than stem fractions with no differences between cattle and sheep. The proportion of acetic acid for cattle (0.75) was higher (P < 0.05) than that for sheep (0.69) while the corresponding propionic acid proportions were lower (P < 0.01) in cattle (0.14) compared with sheep (0.19). These differences were associated with a higher passage rate constant for water flowing from the rumen and a higher digestion rate constant in sheep. It was concluded that both cattle and sheep ate more of the leaf than the stem fraction of the legume Lablab purpureus and that the higher intake of leaf was associated with the shorter time that it was retained in the rumen and was not due to differences in digestibility. Cattle had a smaller relative voluntary intake of legume than sheep, a difference that was associated with the longer time that digesta was retained in the rumen.



1967 ◽  
Vol 21 (3) ◽  
pp. 587-597 ◽  
Author(s):  
D. J. Minson

1. Experiments were conducted with wether sheep in 1964 and 1965 fed pelleted or chopped mature Digitaria decubemes Stent (pangola grass) bay containing different crude protein contents. The voluntary intake of food, digestibility of dry matter and nitrogen, nitrogen retention and apparent time of retention of feed organic matter in the reticulo-rumen were measured. The different crude protein contents of the grass were obtained by applications of urea to the sward 14 and 28 days before cutting for hay in 1964 and 1965 respectively. This treatment increased the crude protein content of the dry matter from 4.9% to 8.7% in 1964 and from 3.7% to 7.2% in 1965. The size of the particles of the ground hay before pelleting is given.2. The mean voluntary intake of chopped fertilized grass was 10% and 54% greater than that of the unfertilized; the voluntary intake of pellets made from fertilized grass was 35% and 75% greater than of those made from the unfertilized grass in 1964 and 1965 respectively.3. The voluntary intake of pellets of unfertilized grass was 7% and 14% greater than that 30% greater than that of chopped fertilized grass in 1964 and 1965 respectively.4. The digestibility of the pellets was less than that of the chopped grass.5. The apparent digestibility of the feed nitrogen was increased by the fertilizer nitrogen, but grinding and pelleting had no consistent effect. Sheep eating chopped or pelleted fertilized hay were in positive nitrogen balance.6. The apparent retention time of organic matter in the reticulo-rumen was longer when the sheep were eating chopped hay than when they were eating pellets. In 1964 the apparent retention time of organic matter in the reticulo-rumen was shorter for chopped and pelleted unfertilized grass than for chopped and pelleted fertilized grass, but in 1965 the order was reversed.7. The relationship between voluntary intake, apparent retention time of organic matter in the rumen and the protein content of the food is discussed.



1981 ◽  
Vol 32 (1) ◽  
pp. 109 ◽  
Author(s):  
DP Poppi ◽  
DJ Minson ◽  
JH Ternouth

Pangola grass (Digitauia decumbens) and Rhodes grass (Chlouis gayana) cut as 6 and 12 week regrowths were separated into leaf and stem fractions and fed ad libitum to four cattle and eight sheep fitted with ruminal cannulae. Cattle and sheep ate more of the leaf than of the stem fraction, and this was associated with the shorter time that neutral detergent fibre (NDF) was retained in the rumeno-reticulum (rumen). To study this difference, measurements were made of the retention time of lignin, passage rate of undigested NDF and water from the rumen, ruminal water volume, potential digestibility of NDF and rate of digestion of NDF by means of the nylon bag technique and in vivo techniques. Possible deficiencies of protein and minerals were minimized from this study by feeding casein and a mineral supplement. Lignin was used to measure the time that indigestible feed fractions remained in the rumen. The retention time of the NDF was closely correlated with the time lignin was retained in the rumen (r = 0.93, P < 0.01). This indicated that the longer retention time and lower intake found with the stem fraction was associated with the slower rate of passage of NDF through the reticulo-omasal orifice. The longer retention time of NDF by cattle compared with sheep could also be explained in this way. The passage rate constant for water leaving the rumen was slightly higher for cattle and sheep fed on leaf fractions and lower for cattle compared to sheep on all diets. The flow of water from the rumen through the reticulo-omasal orifice was 25 and 22% (P < 0.05) more for cattle and sheep respectively when they consumed leaf compared to when they consumed stem. The concentration of NDF in water passing through the reticulo-omasal orifice was similar for leaf and stem fractions (26 and 27 gl-1. There was a higher frequency of ventral ruminal sac contractions in sheep fed on leaf compared with those eating stem. The potential digestibility of the NDF was higher for leaf than stem fractions (0.658 v. 0.600, P < 0.05). Rates of digestion of leaf and stem samples in nylon bags suspended in the rumen of cattle were similar. Samples that had been masticated had a higher digestion rate constant than chopped grass (0.0216 v. 0.0159, P < 0.05). Digestion rate constants calculated from a rumen model showed large differences both between leaf and stem (0.0488 c. 0.0305, P < 0.05) and between cattle and sheep (0.0307 v. 0.0486, P < 0.05). It was suggested that the higher digestion rate constant calculated from a rumen model compared with the nylon bag method was due to additional mastication during rumination. It was concluded that the higher voluntary intake and shorter retention time in the rumen of the leaf than of the stem fraction of grasses was associated with an apparent higher rate of digestion of NDF in vivo, the higher rate of passage of the NDF from the rumen and the higher potential digestibility of the leaf. Sheep were found to retain NDF in the rumen for a shorter time than cattle, and this difference also was associated with a higher rate of digestion of the NDF and more rapid rate of passage of the NDF.



1983 ◽  
Vol 49 (3) ◽  
pp. 463-473 ◽  
Author(s):  
R. M. Dixon ◽  
J. J. Kennelly ◽  
L. P. Milligan

1. Two rumen cannulated steers consuming 5·5 kg air-dry lucerne (Medicago sativa) hay/d given at two-hourly intervals were used to study the kinetics in the rumen of the two particulate markers, 103Ru-labelled Tris-(1,10-phenanthroline) ruthenium II chloride ([103Ru]P) and dysprosium (Dy). Provision of markers was achieved by allowing the steers to eat separatedstems of the hay on to which had been sprayed solutions of the markers.2. The intake of large-particle (retained by a 3·2 mm mesh screen) dry matter (DM) in boluses and the rumen large-particle pool size measured by emptying the rumen wereused to calculate the turnover rate-constant of this pool (1·02 and 1·19/d for steers A and B respectively).3. The decline with time of both [103Ru]P and Dy associated with large-particle DM in raft digesta was best described by two-compartmental kinetics. The first compartment apparently reflected a combination of the processes of mixing of labelled particles throughout the rumen contents, physical migration of marker from the labelled hay, and physical breakdown of large particles to small particles.4. The disappearance with time of [103Ru]P and Dy associated with small particles from the raft, ventral digesta and faeces from 3 to 8 d was well described by a single kinetic compartment with a rate constant similar to that of the second compartment of the large particles from the raft.5. [103Ru]P was rapidly distributed through both the raft and ventral digesta of the rumen. This observation, taken together with measurements of migration of [103Ru]P, suggested that on entering the rumen much of the [103Ru]P did not remain associated with the original feed material.6. The majority (63–64%) of Dy entered the raft digesta and mixed only slowly through the rumen contents. Some Dy (18–27%) apparently migrated rapidly from large particles and to small particles immediately after ingestion and there was also evidence for some slow migration from small to large particles.7. When used under the conditions described for this experiment neither [103Ru]P nor Dy was satisfactory as a marker to trace the passage through the rumen of a particular meal.



1985 ◽  
Vol 105 (1) ◽  
pp. 9-14 ◽  
Author(s):  
D. P. Poppi ◽  
R. E. Hendricksen ◽  
D. J. Minson

SUMMARYIn a study of the effect of animal species on the threshold particle size leaving the rumen, two grasses cut at two stages of growth and one mature legume were separated into leaf and stem fractions and fed to cattle and sheep. Samples of rumen digesta and faeces were used to determine the validity of using a 1·18 mm porosity screen to separate the rumen particles into large and small pools when studying escape of particles from the rumen. Samples of rumen digesta and faeces were collected for the determination of particle size by wet sieving and the calculation of resistance of particles to passage from the rumen relative to small particles retained on a 0·15 mm sieve.Particles < 1·18 mm but > 0·5 mm had a mean relative resistance to passage of 2·0 and 2·6 for cattle and sheep respectively, compared with resistance values of between 10·9 and 31·2 for particles between 1·18 and 2·36 mm. It is suggested that there is no justification for using different threshold particle sizes for sheep and cattle and that a 1·18 mm sieve may be used to divide the rumen contents of both cattle and sheep into two pools of particles with high and low relative resistance to passage from the rumen.



1989 ◽  
Vol 43 (5) ◽  
pp. 791-794 ◽  
Author(s):  
R. O. Carter ◽  
K. R. Carduner ◽  
M. C. Paputa Peck ◽  
D. H. Motry

An explanation is presented for the broadening of intense spectral features observed in the infrared spectra of polyethylene terephthalate fibers prepared as KBr pellets. Since band broadening occurs in the spectra of stronger, undegraded fiber, it is proposed that, the stronger the fiber, the more the difficulty one has in pulverizing the sample. Larger particles result from undegraded polyester, which lead to broader bands for intense features. The local sample pathlength in the region of the large particles is greater than that for small particles, which can be more homogeneously mixed. Fiber samples can also be presented to an infrared microscope either “as is” or flattened. Striking differences exist between spectra for flattened and round fibers that were otherwise similar.



2008 ◽  
Vol 19 (1) ◽  
pp. 61-86 ◽  
Author(s):  
M. SHEARER ◽  
J. M. N. T. GRAY ◽  
A. R. THORNTON

Dense, dry granular avalanches are very efficient at sorting the larger particles towards the free surface of the flow, and finer grains towards the base, through the combined processes of kinetic sieving and squeeze expulsion. This generates an inversely graded particle-size distribution, which is fundamental to a variety of pattern formation mechanisms, as well as subtle size-mobility feedback effects, leading to the formation of coarse-grained lateral levees that create channels in geophysical flows, enhancing their run-out. In this paper we investigate some of the properties of a recent model [Gray, J. M. N. T. & Thornton, A. R. (2005) A theory for particle size segregation in shallow granular free-surface flows. Proc. R. Soc. 461, 1447–1473]; [Thornton, A. R., Gray, J. M. N. T. & Hogg, A. J. (2006) A three-phase mixture theory for particle size segregation in shallow granular free-surface flows. J. Fluid. Mech. 550, 1–25] for the segregation of particles of two sizes but the same density in a shear flow typical of shallow avalanches. The model is a scalar conservation law in space and time, for the volume fraction of smaller particles, with non-constant coefficients depending on depth within the avalanche. It is proved that for steady flow from an inlet, complete segregation occurs beyond a certain finite distance down the slope, no matter what the mixture at the inlet. In time-dependent flow, dynamic shock waves can develop; they are interfaces separating different mixes of particles. Shock waves are shown to be stable if and only if there is a greater concentration of large particles above the interface than below. Constructions with shocks and rarefaction waves are demonstrated on a pair of physically relevant initial boundary value problems, in which a region of all small particles is penetrated from the inlet by either a uniform mixture of particles or by a layer of small particles over a layer of large particles. In both cases, and under a linear shear flow, solutions are constructed for all time and shown to have similar structure for all choices of parameters.



Author(s):  
Liu-xi Cai ◽  
Shun-sen Wang ◽  
Lei Zhang ◽  
Jing-ru Mao ◽  
Zhen-ping Feng ◽  
...  

Reducing the solid particle erosion (SPE) of blades is one of the most important problems for high-parameter steam turbine. Based on the erosion rate models and the particle rebound models of blade materials obtained through the accelerated erosion test under high temperature, a lot of three dimension numerical simulations were conducted in this paper. The influence of particle size on the impingement point distribution on the nozzle surface and the erosion characteristics of control stage nozzle in a supercritical steam turbine were analyzed quantitatively. The size range of the oxide scale particles participating in the erosion damage were extended to 500μm, and some special anti-erosion measures corresponding to different size particles were proposed to reduce the erosion of nozzle. Results show that the erosion of pressure surface in the trailing edge of nozzle is mainly from the high-intensity impingement of particles smaller than 160μm, especially those with the size range from 20μm to 60μm. For the impingement of these small particles, optimizing the profile and cascade structure as well as coating the hard coating on the surface of nozzle can improve the erosion resistance of nozzle. However, these small particles do not impinge the nozzle trailing edge suction surface. The severe erosion damage of suction surface of nozzle was from the impingement again of the particles with the size range from 200μm to 500μm after rebounding from nozzle pressure surface. It is very difficult to resist the erosion damage of these large particles for the hard coating, and separating large particles from main steam before entering the nozzle chamber should be a good choice for improving the erosion resistance of turbine. These studies will provide a technical basis for selecting anti-erosion measures of control stage nozzle.



1974 ◽  
Vol 82 (3) ◽  
pp. 419-422 ◽  
Author(s):  
M. C. Rees ◽  
D. J. Minson ◽  
F. W. Smith

SUMMARYPangola grass grown with and without sulphur fertilizer was cut as a 7-week regrowth to measure voluntary intake and digestibility by sheep. To determine the extent of a simple sulphur deficiency half the sheep on each feed were supplemented with 0·6 gsulphur daily. Retention time of feed in the reticulo-rumen and proportion of feed digested anterior to the duodenum was also determined.Sulphur fertilizer increased the sulphur content of the pasture from 0·09 to 0·15%, voluntary intake of dry matter from 44·4 to 64·1 g/kg W0·75/day and dry-matter digestibility from 55·2 to 60·2%. Sulphur fertilizer reduced the retention time in the reticulo-rumen by 16% and increased the extent of dry-matter digestion before the duodenum from 13 to 34%.Feeding a sulphur supplement increased dry-matter digestibility of the low sulphur feed to that found with sulphur fertilized pangola grass but only accounted for 63% of the difference in voluntary intake.It was concluded that fertilizer sulphur leads to large increases in both voluntary intake and digestibility of sulphur deficient grass and that the improvement in nutritive value may be more than can be achieved by feeding a sulphur supplement.



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