scholarly journals Evaluating the impact of stand composition and competition on white spruce and aspen wood attributes in the boreal mixedwood

2021 ◽  
Author(s):  
◽  
Ryan Jackalin

I evaluated how intra- and inter-specific competition affects the development of eleven wood attributes of trembling aspen (Populus tremuloides Michx.) and white spruce (Picea glauca (Moench) Voss) over 34 years. My analysis was conducted in a mixedwood trial site in Northern British Columbia, Canada, that included treatments consisting of 0, 1000, 2000, 5000, and 10000 stems per hectare of aspen. Competition was found to negatively influence wood attribute development in aspen and positively impact spruce (at low levels of competition). Plot level competition indices were the best predictor of variation in aspen wood attributes, while stand level competition (population density) best explained the majority of spruce wood attributes. Maintaining aspen at lower densities in intimate mixture can have a positive effect on spruce wood quality, while incurring relatively small reductions in spruce volume production and also retaining the ecological benefits associated with managing for mixed stands.

2011 ◽  
Vol 26 (2) ◽  
pp. 82-90 ◽  
Author(s):  
Richard D. Kabzems ◽  
George Harper ◽  
Peter Fielder

Abstract Managing boreal mixed stands of trembling aspen (Populus tremuloides Michx.) and white spruce (Picea glauca [Moench] Voss) is more likely to sustain a diversity of values and has the potential to increase productivity at both the site and landscape levels compared with pure broadleaf or conifer management. In this study, we examine growth of white spruce and aspen after 11 growing seasons over a range of aspen densities created by spot and broadcast treatment of broadleaves using manual and chemical means, aspen spacing, and an untreated control. Results indicate that survival and growth of both spruce and aspen were similar across the range of treatments. Spruce groundline diameter was greater, and height to groundline diameter ratio was lower, for the treatments in which aspen was chemically controlled or uniformly spaced compared with the control. Light measurements at the individual tree level suggested that increased light availability improved white spruce diameter growth. Spruce height growth did not vary by treatment. The status of these experimental mixedwoods was compared with current conifer and mixedwood regeneration evaluations, as well as the preharvest composition of the original stand. After 11 growing seasons, growth of aspen and white spruce indicated that opportunities exist to further modify aspen density to enhance treatment longevity and effectiveness to produce a greater range of boreal mixedwood stand types.


2015 ◽  
Vol 45 (1) ◽  
pp. 78-86 ◽  
Author(s):  
Ya-Lin Hu ◽  
Yue Hu ◽  
De-Hui Zeng ◽  
Xiao Tan ◽  
Scott X. Chang

Exponential fertilization has been shown to be a useful technique for improving seedling quality during nursery production. In this study, we evaluated (i) the impact of exponential vs. conventional fertilization on trembling aspen (Populus tremuloides Michx.) and white spruce (Picea glauca (Moench) Voss) seedlings during nursery production and (ii) the growth performance and nitrogen (N) utilization of those seedlings in response to foxtail barley (Hordeum jubatum L.) competition after the transplantation of seedlings in a greenhouse experiment. Exponential fertilization with an application rate higher than the conventional fertilization increased the internal N reserve in trembling aspen and white spruce seedlings during nursery production and also increased new leaf, new stem, and old stem biomass and N retranslocation rates in trembling aspen seedlings but not in white spruce seedlings after transplantation. Foxtail barley competition decreased N concentrations in seedlings and soil N uptake by the seedlings; however, increased N retranslocation rates with foxtail barley competition were observed in white spruce seedlings but not in trembling aspen seedlings. Our results suggest that the growth performance of seedlings was improved by N loading, whereas the impact of vegetation management was species specific.


2004 ◽  
Vol 34 (2) ◽  
pp. 384-395 ◽  
Author(s):  
Richard Kabzems ◽  
Oscar García

The trembling aspen (Populus tremuloides Michx.) – white spruce (Picea glauca (Moench) Voss) mixed woods near Fort Nelson are distinguished by the large size of individual trees, longevity, and the low occurrence of internal decay in trembling aspen. The development of these forest ecosystems has had limited documentation and may be significantly different than those described in other portions of the boreal forest. At five study stands, stem analysis techniques were used to examine the patterns of height and radial growth over time according to species and structure type. There were two patterns of species establishment that were consistent with the stand structure. In codominant stands, recruitment periods for trembling aspen and white spruce overlapped. The stratified stands were consistently associated with a 29- to 58-year lag in white spruce recruitment. Spruce that were codominant with aspen at the time of sampling had sustained periods of rapid height and diameter growth. White spruce that were later to establish on site had slower rates of height and diameter growth. White spruce ages indicated that a dominant recruitment episode was more common than continuous recruitment. Height and diameter growth of trembling aspen were similar in both stand types. The differences in trembling aspen growth patterns between stands were due to site quality. The white spruce in codominant stands did not appear to go through a period of suppression and then release associated with stand-level trembling aspen mortality, as commonly described for other boreal mixedwoods. The vigor and longevity of trembling aspen in Fort Nelson appear to prolong the period of trembling aspen domination of mixed stands well beyond the time periods observed in other boreal ecosystems.


2007 ◽  
Vol 22 (3) ◽  
pp. 163-170 ◽  
Author(s):  
Ryan J. Klos ◽  
G. Geoff Wang ◽  
Qing-Lai Dang ◽  
Ed W. East

Abstract Kozak's variable exponent taper equation was fitted for balsam poplar (Populus balsamifera L.), trembling aspen (Populus tremuloides Michx.), white spruce (Picea glauca [Moench] Voss), black spruce (Picea mariana [Mill.] B.S.P.), and jack pine (Pinus banksiana Lamb.) in Manitoba. Stem taper variability between two ecozones (i.e., Boreal Shield and Boreal Plains) were tested using the F-test. Regional differences were observed for trembling aspen, white spruce, and jack pine, and for those species, separate ecozone-specific taper equations were developed. However, the gross total volume estimates using the ecozone-specific equations were different from those of the provincial equations by only 2 percent. Although the regional difference in stem form was marginal within a province, a difference of approximately 7 percent of gross total volume estimation was found when our provincial taper equations were compared with those developed in Alberta and Saskatchewan. These results suggest that stem form variation increases with spatial scale and that a single taper equation for each species may be sufficient for each province.


2014 ◽  
Vol 44 (2) ◽  
pp. 162-171 ◽  
Author(s):  
Derek F. Sattler ◽  
Philip G. Comeau ◽  
Alexis Achim

Radial patterns of modulus of elasticity (MOE) were examined for white spruce (Picea glauca (Moench) Voss) and trembling aspen (Populus tremuoides Michx.) from 19 mature, uneven-aged stands in the boreal mixedwood region of northern Alberta, Canada. The main objectives were to (1) evaluate the relationship between pith-to-bark changes in MOE and cambial age or distance from pith; (2) develop species-specific models to predict pith-to-bark changes in MOE; and (3) to test the influences of radial growth, relative vertical height, and tree slenderness (tree height/DBH) on MOE. For both species, cambial age was selected as the best explanatory variable with which to build pith-to-bark models of MOE. For white spruce and trembling aspen, the final nonlinear mixed-effect models indicated that an augmented rate of increase in MOE occurred with increasing vertical position within the tree. For white spruce trees, radial growth and slenderness were found to positively influence maximum estimated MOE. For trembling aspen, there was no apparent effect of vertical position or radial growth on maximum MOE. The results shed light on potential drivers of radial patterns of MOE and will be useful in guiding silvicultural prescriptions.


1999 ◽  
Vol 75 (3) ◽  
pp. 505-513 ◽  
Author(s):  
Rongzhou Man ◽  
Victor J. Lieffers

In boreal mixedwood forests, aspen (Populus tremuloides) and white spruce (Picea glauca) commonly grow in mixture. These species may avoid competition through differential shade tolerance, physical separation of canopies, phenological differences, successional separation, and differences in soil resource utilization. Aspen may also be able to positively affect the growth of white spruce by improving litter decomposition and nutrient cycling rates, controlling grass and shrub competition, ameliorating environmental extremes, and reducing pest attack. These positive relationships likely make mixed-species stands more productive than pure stands of the same species. The evidence regarding the productivity of pure versus mixed aspen/white spruce stands in natural unmanaged forests is examined in this paper. Key words: Tree mixture; productivity; boreal mixedwoods; aspen; white spruce


1959 ◽  
Vol 37 (1) ◽  
pp. 29-34 ◽  
Author(s):  
G. A. Adams

Acid hydrolysis of extractive-free white spruce wood produced a number of neutral and acidic sugars and oligosaccharides. The acidic components were isolated and three of these were shown to be 4-O-methyl-D-glucuronic acid, 2-O-(4-O-methyl-α-D-glucopyranosyluronic acid)-D-xylose, and tentatively O-(4-O-methyl-α-D-glucopyranosyluronic acid)-(1→ 2)-O-β-D-xylopyranosyl-(1→ 4)-D-xylopyranose.


2006 ◽  
Vol 36 (7) ◽  
pp. 1818-1833 ◽  
Author(s):  
Daniel A MacIsaac ◽  
Philip G Comeau ◽  
S Ellen Macdonald

This study assessed the dynamics of gap development in postharvest regeneration in five stands in northwestern Alberta dominated by trembling aspen (Populus tremuloides Michx.). The pattern of gap development over time was determined from analysis of air photographs taken preharvest and 1, 4, 10, and 12 years postharvest. The area of each stand covered by gaps increased after harvest because of the addition of harvest-related gaps over and above those that had been present prior to harvest. The blocks we studied had a combined gap area of up to 29% of stand area 12 years postharvest. We measured regeneration characteristics, microsite, soil, light, and browse conditions in 30 aspen regeneration gaps (gaps in regeneration that were not gaps preharvest and were not due to obvious harvest-related disturbance) 14 years following harvest. Although deciduous trees within postharvest regeneration gaps were the same age as those outside (i.e., in a fully stocked matrix of newly established even-aged aspen stems), they were often suppressed, with significantly lower density and growth. Within the 14-year-old postharvest regenerating aspen stands, aspen height varied from 1 to 11 m; this substantial variability appeared to be largely due to the influence of browsing. There was little evidence of ongoing regeneration within postharvest regeneration gaps, indicating that these gaps will probably persist over time. This may impact future deciduous stocking and volume. It is unknown what may have initiated the formation of these gaps, although results suggest that they are not due to edaphic conditions or disease in the preharvest stands. There is evidence that bluejoint (Calamagrostis canadensis (Michx.) Beauv.) cover and browsing are important factors in the maintenance of postharvest regeneration gaps. The spatial heterogeneity resulting from gaps could be advantageous, however, either as part of ecosystem-based management emulating natural disturbance or as a template for mixedwood management, where white spruce (Picea glauca (Moench) Voss) are established in gaps.


2002 ◽  
Vol 32 (6) ◽  
pp. 1071-1079 ◽  
Author(s):  
Clive Welham ◽  
Brad Seely ◽  
Hamish Kimmins

The ecosystem model FORECAST was used to simulate the yield potential in Saskatchewan mixedwoods of the two-pass harvesting system. The simulated two-pass stand consisted of an overstory population of pure trembling aspen (Populus tremuloides Michx.) with a white spruce (Picea glauca (Moench) Voss) understory. The aspen was removed at year 60, and yields of the understory spruce and resprouting aspen were simulated for 80 years thereafter. The two-pass simulations were compared with two simulated conventional harvesting systems. The first system consisted of a single final harvest at year 140. In the second system, a clearcut was conducted at year 60. White spruce was then planted in the subsequent year at 400, 600, or 800 stems/ha, and aspen also permitted to resprout. Growth was then simulated for a further 80 years. FORECAST projections indicated that the two-pass system might be effective for releasing the white spruce understory, achieving at least a twofold gain in spruce volume relative to conventional methods. Furthermore, total volumes exceeded those derived from the unmanaged stand, while second rotation yields of aspen declined with spruce understory density. These simulations suggest the two-pass harvesting system has strong potential as a tool for mixedwood management.


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