Variability in forest floor at different spatial scales in a natural forest in the Carpathians: effect of windthrows and mesorelief
Spatial variability of humus properties in a natural fir–beech forest was studied along with the influence of windthrows and mesorelief on this variability. In 1720 windthrows the thickness and form of the organic horizons were studied in three positions — mound, pit, and undisturbed control. On undisturbed sites, substantial variability of thickness and forms of the organic horizons was found on a fine scale (0–10 m). Close spatial dependence of some humus characteristics was found on a coarser scale (20–120 m). The mesorelief was found to be one of the key autocorrelation factors. The level of spatial dependence was not uniform; it differed between the fermented and humification horizons and among their forms. The presence of windthrows increased the variability of humus thickness on both fine (0–10 m) and coarse (level of entire locality, i.e., 11 ha) scales. However, windthrows did not increase the variability of organic horizon forms (OHFs) on a fine scale. High variability of OHFs is probably a property of fully developed mature humus in a natural fir–beech forest. On a coarse scale, the presence of pits increased the frequency of fermented zoogenous and humification residual horizons on the study area.