Canopy and emergent white spruce in “pure” broadleaf stands: frequency, predictive models, and ecological importance

2009 ◽  
Vol 39 (10) ◽  
pp. 1997-2004 ◽  
Author(s):  
Steve Cumming ◽  
Mariana Trindade ◽  
David Greene ◽  
S. Ellen Macdonald
Keyword(s):  
Picea Glauca ◽  
Boreal Forests ◽  
Stand Density ◽  
White Spruce ◽  
Stand Age ◽  
Large White ◽  
Forest Management Planning ◽  
Mixedwood Forests ◽  
Seed Sources ◽  
Ecological Importance

In mixedwood boreal forests of western Canada, stands classified as “pure deciduous” by forest inventories sometimes contain a few large white spruce (Picea glauca (Moench) Voss) trees among or emerging from the canopy. These trees are important as regeneration seed sources and for habitat structure. Neither their abundance nor the characteristics of stands in which they occur have previously been quantified. Of 275 “pure aspen” stands in northeastern Alberta, 19.6% contained at least one such spruce detectable in an aerial photograph. These trees were found in stands across the range of sampled canopy heights, densities, age classes, and stand sizes and were often present in the interior of stands, not just on the perimeter. The frequency of 3 ha cells containing at least one spruce was related to (i) stand shape and size, (ii) amount of mature white spruce in adjacent forest, (iii) canopy height, (iv) stand age, and (v) stand density. We conclude that such trees are relatively abundant and widely distributed within the boreal mixedwood forests of Alberta. There is presently no provision to maintain this landscape element within managed forests. More information is needed to determine if or how they should be considered in forest management planning.

scholarly journals Semi-natural and intensive silvicultural systems for the boreal mixedwood forest

1996 ◽  
Vol 72 (3) ◽  
pp. 286-292 ◽  
Author(s):  
V. J. Lieffers ◽  
J. D. Stewart ◽  
R. B. Macmillan ◽  
D. Macpherson ◽  
K. Branter
Keyword(s):  
Populus Tremuloides ◽  
Land Tenure ◽  
Picea Glauca ◽  
Abies Balsamea ◽  
White Spruce ◽  
Growth And Yield ◽  
Mixedwood Forests ◽  
Seed Sources ◽  
Starting Point ◽  
Western Boreal Forest

Boreal mixedwood forests of aspen (Populus tremuloides) and white spruce (Picea glauca) are found on mesic sites in the western boreal forest. In the natural development of mixedwood stands, aspen is usually the first species to dominate the site. However, depending upon spruce seed sources and seedbeds, spruce can establish immediately after disturbance or in the next several decades. In most cases, spruce grow in the understory of deciduous species during its early development. If there are no spruce seed sources, aspen may be the sole tree species for a long period. In most circumstances, however, the longer-lived and taller white spruce eventually becomes the dominant species. If stands remain undisturbed for long periods, they will likely become uneven-aged mixtures of spruce and balsam fir (Abies balsamea). We propose silvicultural systems that will develop stands of a range of compositions, structures and value. As a starting point, we identify eight different mixed-wood compositions that might be identified in stand inventories, and propose various silvicultural treatments, including underplanting of white spruce, understory protection, shelterwood, and uneven-aged management. Fundamental changes in land tenure and silvicultural regulations, and improvements in estimation of growth and yield will be required before this range of management of mixed-woods can be implemented. Key words: aspen, white spruce, shelterwood, Populus tremuloides, Picea glauca, succession, ecosystem management

Are mixtures of aspen and white spruce more productive than single species stands?

1999 ◽  
Vol 75 (3) ◽  
pp. 505-513 ◽  
Author(s):  
Rongzhou Man ◽  
Victor J. Lieffers
Keyword(s):  
Populus Tremuloides ◽  
Picea Glauca ◽  
White Spruce ◽  
Single Species ◽  
Physical Separation ◽  
Soil Resource ◽  
Mixedwood Forests ◽  
Environmental Extremes ◽  
Spruce Stands ◽  
Boreal Mixedwoods

In boreal mixedwood forests, aspen (Populus tremuloides) and white spruce (Picea glauca) commonly grow in mixture. These species may avoid competition through differential shade tolerance, physical separation of canopies, phenological differences, successional separation, and differences in soil resource utilization. Aspen may also be able to positively affect the growth of white spruce by improving litter decomposition and nutrient cycling rates, controlling grass and shrub competition, ameliorating environmental extremes, and reducing pest attack. These positive relationships likely make mixed-species stands more productive than pure stands of the same species. The evidence regarding the productivity of pure versus mixed aspen/white spruce stands in natural unmanaged forests is examined in this paper. Key words: Tree mixture; productivity; boreal mixedwoods; aspen; white spruce

Patterns of initial versus delayed regeneration of white spruce in boreal mixedwood succession

2006 ◽  
Vol 36 (6) ◽  
pp. 1597-1609 ◽  
Author(s):  
Vernon S Peters ◽  
S Ellen Macdonald ◽  
Mark RT Dale
Keyword(s):  
Populus Tremuloides ◽  
Picea Glauca ◽  
Fire Severity ◽  
Mineral Soil ◽  
White Spruce ◽  
Mixedwood Forests ◽  
Regeneration Period ◽  
Postfire Regeneration ◽  
Distance To Seed Source ◽  
Mixedwood Stands

The timing of white spruce regeneration in aspen (Populus tremuloides Michx.) – white spruce (Picea glauca (Moench) Voss) boreal mixedwood stands is an important factor in stand development. We examined boreal mixedwood stands representing a 59-year period of time since fire and determined (1) whether and when a delayed regeneration period of white spruce occurred, (2) whether the relative abundance of initial (<20 years) versus delayed (≥20 years postfire) regeneration is related to seed availability at the time of the fire, and (3) what are the important regeneration substrates for initial versus delayed regeneration. Initial regeneration occurred primarily on mineral soil or humus, while delayed regeneration established primarily on logs and peaked 38–44 years after fire. Of the 20 stands investigated, seven were dominated by initial regeneration, six were dominated by delayed regeneration, and seven were even mixtures of both. The dominance of a site by initial or delayed regeneration could not be simply explained by burn timing relative to mast years or distance to seed source; our results suggested that fire severity and the competitive influence of initial regeneration on delayed regeneration were important at fine scales. Based on our results we describe several possible postfire successional pathways for boreal mixedwood forests.

Decadal observations of tree regeneration following fire in boreal forests

2004 ◽  
Vol 34 (2) ◽  
pp. 267-273 ◽  
Author(s):  
Jill F Johnstone ◽  
F S Chapin III ◽  
J Foote ◽  
S Kemmett ◽  
K Price ◽  
...  
Keyword(s):  
British Columbia ◽  
Populus Tremuloides ◽  
Picea Glauca ◽  
Boreal Forests ◽  
Pinus Contorta ◽  
Stand Structure ◽  
Stand Density ◽  
Tree Regeneration ◽  
Direct Observations ◽  
Highly Correlated

This paper presents data on early postfire tree regeneration. The data were obtained from repeated observations of recently burned forest stands along the Yukon – British Columbia border and in interior Alaska. Postfire measurements of tree density were made periodically for 20–30 years, providing direct observations of early establishment patterns in boreal forest. Recruitment rates of the dominant tree species in both study areas were highest in the first 5 years after fire, and additional net establishment was not observed after 10 years. The postfire population of spruce (Picea mariana (Mill.) BSP and Picea glauca (Moench) Voss s.l.) remained constant after the first decade in the two study areas. Populations of aspen (Populus tremuloides Michx.) and lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) both declined after 10 years in mixed-species stands along the Yukon – British Columbia border. Mortality rates of aspen and pine were positively correlated with their initial densities, indicating that thinning occurred as a density-dependent process. At all sites, measurements of stand density and composition made early were highly correlated with those made late in the monitoring period, indicating that patterns of stand structure initiated within a few years after fire are maintained through subsequent decades of stand development.

Terminal leader failure in white spruce plantations in northern Minnesota

1986 ◽  
Vol 16 (3) ◽  
pp. 648-650 ◽  
Author(s):  
R. Bruce Harding
Keyword(s):  
White Spruce ◽  
Frost Damage ◽  
Height Growth ◽  
Size Class ◽  
Site Quality ◽  
Stand Age ◽  
Growth Reduction ◽  
Seed Sources ◽  
Terminal Bud ◽  
Replacement Process

Terminal leader failure from bud mortality occurred in 46% of plantation white spruce (Piceaglauca (Moench) Voss) trees, 19–43 years old. Frost damage was identified as the causal factor of bud mortality, with all plantations being established from seed sources of local origin. Large differences in terminal bud failure rates were noted between year of observation, tree size class, and stand age-class. Dominant trees, 33–43 years old, had a 3 year failure rate ranging from 16 to 51 %. For dominant trees, an 18% (9 cm) height growth reduction per year of incidence was noted. The lateral replacement process could account for greater than a 2 m height reduction for dominants at age 50 years, confounding estimates of potential site quality. The height growth of overtopped trees responded differently to frost damage than dominants, with replacement laterals averaging 38% (8 cm) greater height growth than apical terminals of undamaged trees.

scholarly journals A partial deciduous canopy, coupled with site preparation, produces excellent growth of planted white spruce

2019 ◽  
Vol 49 (3) ◽  
pp. 270-280 ◽  
Author(s):  
Victor J. Lieffers ◽  
Derek Sidders ◽  
Tim Keddy ◽  
Kevin A. Solarik ◽  
Peter Blenis
Keyword(s):  
Picea Glauca ◽  
Boreal Forests ◽  
High Speed ◽  
White Spruce ◽  
Site Preparation ◽  
Stem Volume ◽  
Soil Mixing ◽  
Excellent Growth ◽  
Total Growth ◽  
Best Estimates

Survival and growth of planted white spruce (Picea glauca (Moench) Voss) were assessed at year 15 in boreal mixedwood stands of northern Alberta, Canada, in stands that were deciduous-dominated prior to logging or were conifer-dominated. Three overstory retention levels (0%, 50%, and 75% retention) and four site preparation treatments (mound, high speed mix, scalp, and no treatment) were evaluated. In deciduous-dominated stands, planted spruce performed best in the 50% retention; here, stem volume was at least double that of any other retention treatment after 15 years. In contrast, spruce had reduced growth in coniferous-dominated stands in both 50% and 75% retention treatments compared with the 0% retention. Survival of planted spruce was unaffected by level of retention, but survival was lower in coniferous-dominated stands than in deciduous-dominated stands; in the coniferous-dominated stands, survival was better with mounding and mixing and lowest with scalp treatments. All height variables tended to be greater in the mix and mound site preparation treatments. Finally, the best estimates of future total growth (regenerated spruce and deciduous combined) in the coniferous-dominated stands were in the clearcut treatment. In terms of regenerated spruce growth, the best estimates occurred in the deciduous-dominated – 50% retention stand planted with soil mixing–mounding treatments, where projected growth of spruce was comparable with that of open-grown and tended stands in Alberta’s boreal forests.

scholarly journals Regeneration of aspen following partial and strip understory protection harvest in boreal mixedwood forests

2009 ◽  
Vol 85 (4) ◽  
pp. 631-638 ◽  
Author(s):  
Alison D Lennie ◽  
Simon M Landhäusser ◽  
Victor J Lieffers ◽  
Derek Sidders
Keyword(s):  
Forest Management ◽  
Key Words ◽  
Populus Tremuloides ◽  
Picea Glauca ◽  
Basal Area ◽  
White Spruce ◽  
Mixedwood Forests ◽  
Partial Harvest ◽  
Second Year

Trembling aspen regeneration was studied in 2 types of partial harvest systems designed to harvest mature aspen but protect immature spruce and encourage natural aspen regeneration. Two partial harvest systems, where the residual aspen was either left in strips or was dispersed uniformly, were compared to traditional clearcuts. After the first and second year since harvest, aspen sucker density and growth was similar between the 2 partial harvests, but was much lower than in the clearcuts. However, in the partial cuts the regeneration density was very much dependent on the location relative to residual trees. The density of regeneration was inversely related to the basal area of residual aspen; however, sucker height was inversely related to the basal area of the residual spruce. Although there were adequate numbers of suckers after partial harvest, their viability and contribution to the long-term productivity of these mixedwood stands is not clear. Key words: silvicultural systems, forest management, residual canopy, white spruce, Populus tremuloides, Picea glauca, traffic

CO2 stimulation and response mechanisms vary with light supply in boreal conifers

2020 ◽  
Author(s):  
Qing-Lai Dang ◽  
Jacob Marfo ◽  
Fengguo Du ◽  
Rongzhou Man ◽  
Sahari Inoue
Keyword(s):  
Elevated Co2 ◽  
Picea Glauca ◽  
Boreal Forests ◽  
Black Spruce ◽  
Net Photosynthesis ◽  
Growing Season ◽  
White Spruce ◽  
Light Supply ◽  
Ecophysiological Responses ◽  
Light Effects

Abstract Aims Black spruce (Picea mariana [Mill.] B.S.P.) and white spruce (Picea glauca [Moench] Voss.) are congeneric species. Both are moderately shade tolerant and widely distributed across North American boreal forests. Methods To understand light effects on their ecophysiological responses to elevated [CO2], 1-year old seedlings were exposed to 360 and 720 µmol mol -1 [CO2] at three light conditions (100, 50 and 30% of full light in the greenhouse). Foliar gas exchanges were measured in the mid- and late-growing season. Important Findings Elevated [CO2] increased net photosynthesis (Pn) and photosynthetic water use efficiency, but it reduced stomatal conductance and transpiration. The stimulation of photosynthesis by CO2 was greatest at 50% light and smallest at 100%. Photosynthesis, maximum carboxylation rate (Vcmax) and light saturated rate of electron transport (Jmax) all decreased with decreasing light. Elevated [CO2] significantly reduced Vcmax across all light treatments and both species in mid-growing season. However, the effect of elevated [CO2] became insignificant at 30% light later in the growing season, with the response being greater in black spruce than in white spruce. Elevated [CO2] also reduced Jmax in white spruce in both measurements while the effect became insignificant at 30% light later in the growing season. However, the effect on black spruce varied with time. Elevated [CO2] reduced Jmax in black spruce in mid-growing season in all light treatments and the effect became insignificant at 30% light later in the growing season, while it increased Jmax later in the season at 100% and 50% light. These results suggest that both species benefited from elevated CO2, and that the responses varied with light supply, such that the response was primarily physiological at 100% and 50% light, while it was primarily morphological at 30% light.

scholarly journals Variation of White Spruce Carbon Content with Age, Height, Social Classes and Silvicultural Management

Energies ◽  
2021 ◽  
Vol 14 (23) ◽  
pp. 8015
Author(s):  
Cyriac S. Mvolo ◽  
James D. Stewart ◽  
Christopher Helmeste ◽  
Ahmed Koubaa
Keyword(s):  
Carbon Content ◽  
Picea Glauca ◽  
Stand Density ◽  
White Spruce ◽  
Carbon Accounting ◽  
Accuracy And Precision ◽  
Freeze Dried ◽  
Stand Density Management ◽  
And Control ◽  
Volatile Carbon

The accuracy and precision with which carbon amounts have been accounted for in forests have been questioned. As countries seek to comply with agreements to reduce global warming and industries seek to maximize bioenergy potential, this matter has increased international concern. White spruce (Picea glauca (Moench) Voss) stand density management trials in the Petawawa Research Forest, Ontario, Canada, were sampled to evaluate carbon concentration variation within trees and plots of differing stand density. Sample-drying methodologies were also tested to compare freeze-dried carbon (FDC) and oven-dried carbon (ODC) measurements. The average FDC was 51.80 ± 1.19%, and the corrected freeze-dried carbon content (FDCCOR) was 51.76 ± 1.33%. The average ODC was 49.10 ± 0.92%, and the average volatile carbon fraction (Cvol) was 2.67 ± 1.71%. FDC was higher than ODC (mean of the differences = 2.52) and generally more variable. ODC significantly decreased radially and longitudinally. FDC was significantly affected by thinning, where heavy treatments resulted in the highest FDC amounts compared to medium, light, and control treatments. In addition to reducing carbon content (CC), drying influences wood CC in many ways that are still to be elucidated. The results of this study suggest that ODC should continue to be used within the bioenergy industry, while FDC must become the preferred standard for carbon accounting protocols.

scholarly journals Predicting white spruce cone crops in the boreal forests of southern and central Yukon

2017 ◽  
Vol 47 (1) ◽  
pp. 47-52 ◽  
Author(s):  
C.J. Krebs ◽  
M. O’Donoghue ◽  
Shawn Taylor ◽  
A.J. Kenney ◽  
E.J. Hofer ◽  
...  
Keyword(s):  
Picea Glauca ◽  
Boreal Forests ◽  
Predictive Performance ◽  
Information Criterion ◽  
White Spruce ◽  
Daily Maximum ◽  
Degree Days ◽  
The Difference ◽  
Maximum Temperatures ◽  
Regional Synchrony

White spruce (Picea glauca (Moench) Voss) cone crops were measured at five regional centers in southern and central Yukon for 30 years at one site from 1986 to 2015 and at four other sites during 9 to 11 years to select the best climatic model that uses cues from growing season temperature and rainfall to predict the size of cone crops. We evaluated six climatic models that use summer temperature and rainfall of years t – 1 and t – 2 to predict cone crops in year t. July temperatures provided the best predictors of white spruce cone crops, and no rainfall variable was related to the size of cone crops. We explored three variants of July temperatures: mean temperature, degree-days > 5 °C, and maximum temperatures. For each of these, we used the ΔT model that uses the difference in the July temperature measures of years t – 1 and t – 2. We compared the resulting six models with corrected Akaike’s information criterion (AICc) to determine their relative predictive performance. The best model combined ΔT measures of degree-days > 5 °C and the four highest daily maximum July temperatures with R2 = 0.65. By comparison, the ΔT model involving only mean July temperatures was less successful (R2 = 0.49). There was good regional synchrony (rp = 0.7 to 0.8) in high cone crops over southern and central Yukon during 1986 to 2015.

Sign in / Sign up

Export Citation Format

Share Document