scholarly journals THE LOCUST TEGULA: SIGNIFICANCE FOR FLIGHT RHYTHM GENERATION, WING MOVEMENT CONTROL AND AERODYNAMIC FORCE PRODUCTION

1993 ◽  
Vol 182 (1) ◽  
pp. 229-253 ◽  
Author(s):  
H Wolf
Keyword(s):  
Flight Control ◽  
Aerodynamic Force ◽  
Motor Pattern ◽  
Vertical Component ◽  
Sense Organ ◽  
Force Production ◽  
Movement Control ◽  
Rhythm Generation ◽  
Wing Movement ◽  
Before And After

The tegula, a complex sense organ associated with the wing base of the locust, plays an important role in the generation of the flight motor pattern. Here its function in the control of wing movement and aerodynamic force production is described.The vertical component of forewing movement was monitored while recording intracellularly from flight motoneurones during stationary flight. First, in accordance with previous electrophysiological results, stimulation of hindwing tegula afferents was found to reset the wingstroke to the elevation phase in a well-coordinated manner. Second, recordings made before and after removal of fore- and hindwing tegulae were compared. This comparison demonstrated that the delayed onset of elevator motoneurone activity caused by tegula removal is accompanied by a corresponding delay in the upstroke movement of the wings.The consequences of this delayed upstroke for aerodynamic force production were investigated by monitoring wing movements and lift generation simultaneously. A marked decrease in net lift generation was observed following tegula removal. Recordings of wing pronation indicate that this decrease in lift is primarily due to the delayed upstroke movement - that is, to a delay of the wings near the aerodynamically unfavourable downstroke position.It is concluded that the tegula of the locust hindwing signals to the nervous system the impending completion of the wing downstroke and allows initiation of the upstroke movement immediately after the wings have reached the lower reversal point of the wingstroke. The functional significance of tegula feedback and central rhythm generation for locust flight control are discussed.

scholarly journals Deformable wing kinematics in the desert locust: how and why do camber, twist and topography vary through the stroke?

2008 ◽  
Vol 6 (38) ◽  
pp. 735-747 ◽  
Author(s):  
Simon M. Walker ◽  
Adrian L. R. Thomas ◽  
Graham K. Taylor
Keyword(s):  
Flight Control ◽  
High Speed ◽  
Schistocerca Gregaria ◽  
Aerodynamic Force ◽  
Angle Of Attack ◽  
Force Production ◽  
Linear Increase ◽  
Trailing Edge ◽  
Wing Kinematics ◽  
Drag Ratio

Here, we present a detailed analysis of the wing kinematics and wing deformations of desert locusts ( Schistocerca gregaria , Forskål) flying tethered in a wind tunnel. We filmed them using four high-speed digital video cameras, and used photogrammetry to reconstruct the motion of more than 100 identified points. Whereas the hindwing motions were highly stereotyped, the forewing motions showed considerable variation, consistent with a role in flight control. Both wings were positively cambered on the downstroke. The hindwing was cambered through an ‘umbrella effect’ whereby the trailing edge tension compressed the radial veins during the downstroke. Hindwing camber was reversed on the upstroke as the wing fan corrugated, reducing the projected area by 30 per cent, and releasing the tension in the trailing edge. Both the wings were strongly twisted from the root to the tip. The linear decrease in incidence along the hindwing on the downstroke precisely counteracts the linear increase in the angle of attack that would otherwise occur in root flapping for an untwisted wing. The consequent near-constant angle of attack is reminiscent of the optimum for a propeller of constant aerofoil section, wherein a linear twist distribution allows each section to operate at the unique angle of attack maximizing the lift to drag ratio. This implies tuning of the structural, morphological and kinematic parameters of the hindwing for efficient aerodynamic force production.

scholarly journals Flight mechanics and control of escape manoeuvres in hummingbirds. II. Aerodynamic force production, flight control and performance limitations

2016 ◽  
Vol 219 (22) ◽  
pp. 3532-3543 ◽  
Author(s):  
Bo Cheng ◽  
Bret W. Tobalske ◽  
Donald R. Powers ◽  
Tyson L. Hedrick ◽  
Yi Wang ◽  
...  
Keyword(s):  
Flight Control ◽  
Aerodynamic Force ◽  
Force Production ◽  
Flight Mechanics ◽  
Performance Limitations ◽  
And Performance ◽  
And Control

scholarly journals The Effects of Spinal Manipulation on Motor Unit Behavior

2021 ◽  
Vol 11 (1) ◽  
pp. 105
Author(s):  
Lucien Robinault ◽  
Aleš Holobar ◽  
Sylvain Crémoux ◽  
Usman Rashid ◽  
Imran Khan Niazi ◽  
...  
Keyword(s):  
Motor Unit ◽  
Conduction Velocity ◽  
Spinal Manipulation ◽  
Maximum Voluntary Contraction ◽  
Force Production ◽  
Movement Control ◽  
Position Sense ◽  
Joint Position Sense ◽  
Voluntary Contraction ◽  
Passive Movement

Over recent years, a growing body of research has highlighted the neural plastic effects of spinal manipulation on the central nervous system. Recently, it has been shown that spinal manipulation improved outcomes, such as maximum voluntary force and limb joint position sense, reflecting improved sensorimotor integration and processing. This study aimed to further evaluate how spinal manipulation can alter neuromuscular activity. High density electromyography (HD sEMG) signals from the tibialis anterior were recorded and decomposed in order to study motor unit changes in 14 subjects following spinal manipulation or a passive movement control session in a crossover study design. Participants were asked to produce ankle dorsiflexion at two force levels, 5% and 10% of maximum voluntary contraction (MVC), following two different patterns of force production (“ramp” and “ramp and maintain”). A significant decrease in the conduction velocity (p = 0.01) was observed during the “ramp and maintain” condition at 5% MVC after spinal manipulation. A decrease in conduction velocity suggests that spinal manipulation alters motor unit recruitment patterns with an increased recruitment of lower threshold, lower twitch torque motor units.

Postactivation Potentiation of Force Is Independent of H-Reflex Excitability

2008 ◽  
Vol 3 (2) ◽  
pp. 219-231 ◽  
Author(s):  
Matthew J. Hodgson ◽  
David Docherty ◽  
E. Paul Zehr
Keyword(s):  
Isometric Force ◽  
Plantar Flexion ◽  
Force Production ◽  
H Reflex ◽  
Postactivation Potentiation ◽  
Neural Excitability ◽  
Reflex Excitability ◽  
Before And After ◽  
History Of ◽  
A Minor

The contractile history of muscle can potentiate electrically evoked force production. A link to voluntary force production, related in part to an increase in reflex excitability, has been suggested.Purpose:Our purpose was to quantify the effect of postactivation potentiation on voluntary force production and spinal H-reflex excitability during explosive plantar fexion actions.Methods:Plantar flexor twitch torque, soleus H-reflex amplitudes, and the rate of force development of explosive plantar fexion were measured before and after 4 separate conditioning trials (3 × 5 s maximal contractions).Results:Twitch torque and rate of force production during voluntary explosive plantar flexion were significantly increased (P < .05) while H-reflex amplitudes remained unchanged. Although twitch torque was significantly higher after conditioning, leading to a small increase in the rate of voluntary force production, this was unrelated to changes in reflex excitability.Conclusion:We conclude that postactivation potentiation may result in a minor increase in the rate of voluntary isometric force production that is unrelated to neural excitability.

scholarly journals A Data Driven Change-point Epidemic Model for Assessing the Impact of Large Gathering and Subsequent Movement Control Order on COVID-19 Spread in Malaysia

2020 ◽  
Author(s):  
Sarat C. Dass ◽  
Wai M. Kwok ◽  
Gavin J. Gibson ◽  
Balvinder S. Gill ◽  
Bala M. Sundram ◽  
...  
Keyword(s):  
Change Point ◽  
Movement Control ◽  
Disease Spread ◽  
Time Period ◽  
Before And After ◽  
Quantitative Understanding ◽  
Control Order ◽  
Short Time ◽  
The Impact ◽  
Second Wave

AbstractThe second wave of COVID-19 in Malaysia is largely attributed to a mass gathering held in Sri Petaling between February 27, 2020 and March 1, 2020, which contributed to an exponential rise of COVID-19 cases in the country. Starting March 18, 2020, the Malaysian government introduced four consecutive phases of a Movement Control Order (MCO) to stem the spread of COVID-19. The MCO was implemented through various non-pharmaceutical interventions (NPIs). The reported number of cases reached its peak by the first week of April and then started to reduce, hence proving the effectiveness of the MCO. To gain a quantitative understanding of the effect of MCO on the dynamics of COVID-19, this paper develops a class of mathematical models to capture the disease spread before and after MCO implementation in Malaysia. A heterogeneous variant of the Susceptible-Exposed-Infected-Recovered (SEIR) model is developed with additional compartments for asymptomatic transmission. Further, a change-point is incorporated to model the before and after disease dynamics, and is inferred based on data. Related statistical analyses for inference are developed in a Bayesian framework and are able to provide quantitative assessments of (1) the impact of the Sri Petaling gathering, and (2) the extent of decreasing transmission during the MCO period. The analysis here also quantitatively demonstrates how quickly transmission rates fall under effective NPI implemention within a short time period.

scholarly journals Effects of Acupuncture on Explosive Force Production by the Healthy Female Shoulder Joint

2020 ◽  
Vol 2020 ◽  
pp. 1-7
Author(s):  
I-Lin Wang ◽  
Yi-Ming Chen ◽  
Jun Wang ◽  
Rui Hu ◽  
Ke-Ke Zhang ◽  
...  
Keyword(s):  
Shoulder Joint ◽  
Force Production ◽  
Maximum Speed ◽  
Maximum Torque ◽  
Shoulder Muscles ◽  
Potentiation Effect ◽  
Average Maximum ◽  
Significant Difference ◽  
Before And After ◽  
Explosive Force

Background. Acupuncture is often used to treat chronic conditions, such as pain. In recent years, given the importance of the explosive forces generated by shoulder muscles for the completion of motor tasks, studies in which nerves were stimulated through acupuncture to increase the explosive forces were conducted. This study explored the effect of acupuncture on explosive force production by the muscles of the female shoulder joint. Methods. Eighteen healthy women underwent shoulder adduction (Add), abduction (Abd), flexion (Flex), and extension (Ext) tests with an isokinetic measurement system. Acupuncture was used to stimulate the Zhongfu (LU1), Tianfu (LI3), Xiabai (LU4), Binao (LI14), Naohui (SJ13), Jianliao (SJ14), and Xiaoluo (SJ12) points, and electromyography (EMG) signals were recorded before and after acupuncture. Results. After acupuncture, there was a significant difference in the average maximum work, the average maximum power, the average maximum speed, the total work in Add/Abd and Flex/Ext, the EMG signals, and the stiffness of the muscles in Abd and Ext ( P < 0.05 ). There were no significant differences in the average maximum torque in Abd or Flex. Conclusion. Based on the results, there may be a significant correlation between the manipulation of different acupoints by acupuncture and the average maximum torque and stiffness. Acupuncture may stimulate nerves to activate muscles and induce a postactivation potentiation effect that improves explosive force production. Therefore, acupuncture as an auxiliary tool may increase the explosive forces generated by acupoint-related muscles by stimulating nerves.

scholarly journals A chordwise offset of the wing-pitch axis enhances rotational aerodynamic forces on insect wings: a numerical study

2019 ◽  
Vol 16 (155) ◽  
pp. 20190118 ◽  
Author(s):  
Wouter G. van Veen ◽  
Johan L. van Leeuwen ◽  
Florian T. Muijres
Keyword(s):  
Flight Control ◽  
Numerical Study ◽  
Force Production ◽  
Aerodynamic Forces ◽  
Rotational Force ◽  
Wing Motion ◽  
Forward Stroke ◽  
Insect Wings ◽  
Force Mechanism ◽  
Numerical Parametric Study

Most flying animals produce aerodynamic forces by flapping their wings back and forth with a complex wingbeat pattern. The fluid dynamics that underlies this motion has been divided into separate aerodynamic mechanisms of which rotational lift, that results from fast wing pitch rotations, is particularly important for flight control and manoeuvrability. This rotational force mechanism has been modelled using Kutta–Joukowski theory, which combines the forward stroke motion of the wing with the fast pitch motion to compute forces. Recent studies, however, suggest that hovering insects can produce rotational forces at stroke reversal, without a forward motion of the wing. We have conducted a broad numerical parametric study over a range of wing morphologies and wing kinematics to show that rotational force production depends on two mechanisms: (i) conventional Kutta–Joukowski-based rotational forces and (ii) a rotational force mechanism that enables insects with an offset of the pitch axis relative to the wing's chordwise symmetry axis to generate rotational forces in the absence of forward wing motion. Because flying animals produce control actions frequently near stroke reversal, this pitch-axis-offset dependent aerodynamic mechanism may be particularly important for understanding control and manoeuvrability in natural flyers.

Deformable Blade Element and Unsteady Vortex Lattice Fluid-Structure Interaction Modeling of a 2D Flapping Wing

2020 ◽  
Author(s):  
Joseph Reade ◽  
Mark A. Jankauski
Keyword(s):  
Vortex Lattice ◽  
Normal Force ◽  
Aerodynamic Force ◽  
Fluid Structure Interaction ◽  
Force Production ◽  
Flapping Wing ◽  
Energetic Efficiency ◽  
Fluid Structure ◽  
Structure Interaction ◽  
Blade Element

Abstract Flapping insect wings experience appreciable deformation due to aerodynamic and inertial forces. This deformation is believed to benefit the insect’s aerodynamic force production as well as energetic efficiency. However, the fluid-structure interaction (FSI) models used to estimate wing deformations are often computationally demanding and are therefore challenged by parametric studies. Here, we develop a simple FSI model of a flapping wing idealized as a two-dimensional pitching-plunging airfoil. Using the Lagrangian formulation, we derive the reduced-order structural framework governing wing’s elastic deformation. We consider two fluid models: quasi-steady Deformable Blade Element Theory (DBET) and Unsteady Vortex Lattice Method (UVLM). DBET is computationally economical but does not provide insight into the flow structure surrounding the wing, whereas UVLM approximates flows but requires more time to solve. For simple flapping kinematics, DBET and UVLM produce similar estimates of the aerodynamic force normal to the surface of a rigid wing. More importantly, when the wing is permitted to deform, DBET and UVLM agree well in predicting wingtip deflection and aerodynamic normal force. The most notable difference between the model predictions is a roughly 20° phase difference in normal force. DBET estimates wing deformation and force production approximately 15 times faster than UVLM for the parameters considered, and both models solve in under a minute when considering 15 flapping periods. Moving forward, we will benchmark both low-order models with respect to high fidelity computational fluid dynamics coupled to finite element analysis, and assess the agreement between DBET and UVLM over a broader range of flapping kinematics.

Roll aerodynamic characteristics study of an unmanned aerial vehicle based on circulation control technology

2017 ◽  
Vol 233 (3) ◽  
pp. 871-882
Author(s):  
Kun Chen ◽  
Zhiwei Shi ◽  
Jiachen Zhu ◽  
Haiyang Wang ◽  
Junquan Fu
Keyword(s):  
Unmanned Aerial Vehicle ◽  
Flight Control ◽  
Attitude Control ◽  
Aerodynamic Force ◽  
Flight Test ◽  
Scale Model ◽  
Maximum Efficiency ◽  
Control Technology ◽  
Circulation Control ◽  
Aerial Vehicle

To explore the control efficiency of circulation flow control technology, a circulation control actuator with an independent gas source has been designed and applied in roll attitude control of a small unmanned aerial vehicle. The circulation control devices are arranged at the two ends of the wing on an unmanned aerial vehicle scale model, the changes in aerodynamic force and aerodynamic moment caused by turning on the actuator are measured in a wind tunnel, and the flow field characteristics are analysed using particle image velocimetry technology. The flight control effect of the roll attitude is verified via a flight test. Experimental and flight test results show that the control of roll attitude can be achieved by turning on the circulation control actuator on one side, and the maximum efficiency that the circulation control generates is equivalent to 8° aileron deflection with production of a favorable yaw moment to achieve a coordinated turn. The circulation control actuator can increase lift and reduce drag when opened on both sides simultaneously. The maximum lift-to-drag ratio of the UAV increased from 5 to 9, and this approach can also suppress flow separation and delay stall at high angles of attack. The aileron or trailing edge flaps can be replaced with circulation control actuators, and the circulation control technology can also be applied to aerodynamic performance improvement and flight control in other types of aircraft.

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