Chapter 4 further develops the theory of the conservation of living systems. Natural resource management problems are analyzed using optimal control methods. Natural resources are the state variables of the problem and management instruments are control variables. Management might include harvest, culling, restocking, reseeding, and replanting, or interventions affecting, for example, the fire regime, hydrological flows, the structure of habitats, the functioning of the system, and the ecosystem processes involved. The chapter considers three types of systems: aquatic systems, forest systems, and rangelands. It shows how the methods developed to model conversion/conservation decisions in all cases embed the Hotelling arbitrage condition. It shows how the level of conservation in each type of system is impacted by access rules, and the array of benefits obtained from the system.