Three-dimensional kinematics and limb kinetic energy of running cockroaches.

1997 ◽  
Vol 200 (13) ◽  
pp. 1919-1929 ◽  
Author(s):  
R Kram ◽  
B Wong ◽  
R J Full
Keyword(s):  
Kinetic Energy ◽  
High Speed ◽  
Mechanical Energy ◽  
Center Of Mass ◽  
Three Dimensional ◽  
The Body ◽  
Morphological Data ◽  
Fast Running ◽  
Total Mechanical Energy ◽  
Reaction Forces

We tested the hypothesis that fast-running hexapeds must generate high levels of kinetic energy to cycle their limbs rapidly compared with bipeds and quadrupeds. We used high-speed video analysis to determine the three-dimensional movements of the limbs and bodies of cockroaches (Blaberus discoidalis) running on a motorized treadmill at 21 cm s-1 using an alternating tripod gait. We combined these kinematic data with morphological data to calculate the mechanical energy produced to move the limbs relative to the overall center of mass and the mechanical energy generated to rotate the body (head + thorax + abdomen) about the overall center of mass. The kinetic energy involved in moving the limbs was 8 microJ stride-1 (a power output of 21 mW kg-1, which was only approximately 13% of the external mechanical energy generated to lift and accelerate the overall center of mass at this speed. Pitch, yaw and roll rotational movements of the body were modest (less than +/- 7 degrees), and the mechanical energy required for these rotations was surprisingly small (1.7 microJ stride-1 for pitch, 0.5 microJ stride-1 for yaw and 0.4 microJ stride-1 for roll) as was the power (4.2, 1.2 and 1.1 mW kg-1, respectively). Compared at the same absolute forward speed, the mass-specific kinetic energy generated by the trotting hexaped to swing its limbs was approximately half of that predicted from data on much larger two- and four-legged animals. Compared at an equivalent speed (mid-trotting speed), limb kinetic energy was a smaller fraction of total mechanical energy for cockroaches than for large bipedal runners and hoppers and for quadrupedal trotters. Cockroaches operate at relatively high stride frequencies, but distribute ground reaction forces over a greater number of relatively small legs. The relatively small leg mass and inertia of hexapeds may allow relatively high leg cycling frequencies without exceptionally high internal mechanical energy generation.

scholarly journals Race Walking Ground Reaction Forces at Increasing Speeds: A Comparison with Walking and Running

Symmetry ◽  
2019 ◽  
Vol 11 (7) ◽  
pp. 873
Author(s):  
Gaspare Pavei ◽  
Dario Cazzola ◽  
Antonio La Torre ◽  
Alberto E. Minetti
Keyword(s):  
High Speed ◽  
Three Dimensional ◽  
The Body ◽  
Ground Reaction Forces ◽  
Centre Of Mass ◽  
Walking Gait ◽  
Wide Range ◽  
Reaction Forces ◽  
Race Walking ◽  
Walking Speeds

Race walking has been theoretically described as a walking gait in which no flight time is allowed and high travelling speed, comparable to running (3.6–4.2 m s−1), is achieved. The aim of this study was to mechanically understand such a “hybrid gait” by analysing the ground reaction forces (GRFs) generated in a wide range of race walking speeds, while comparing them to running and walking. Fifteen athletes race-walked on an instrumented walkway (4 m) and three-dimensional GRFs were recorded at 1000 Hz. Subjects were asked to performed three self-selected speeds corresponding to a low, medium and high speed. Peak forces increased with speeds and medio-lateral and braking peaks were higher than in walking and running, whereas the vertical peaks were higher than walking but lower than running. Vertical GRF traces showed two characteristic patterns: one resembling the “M-shape” of walking and the second characterised by a first peak and a subsequent plateau. These different patterns were not related to the athletes’ performance level. The analysis of the body centre of mass trajectory, which reaches its vertical minimum at mid-stance, showed that race walking should be considered a bouncing gait regardless of the presence or absence of a flight phase.

Force platforms as ergometers

1975 ◽  
Vol 39 (1) ◽  
pp. 174-179 ◽  
Author(s):  
G. A. Cavagna
Keyword(s):  
Body Weight ◽  
Mechanical Energy ◽  
Center Of Mass ◽  
Force Plate ◽  
Vertical Displacement ◽  
The Body ◽  
Mechanical Work ◽  
Vertical Force ◽  
External Work ◽  
Total Mechanical Energy

Walking and running on the level involves external mechanical work, even when speed averaged over a complete stride remains constant. This work must be performed by the muscles to accelerate and/or raise the center of mass of the body during parts of the stride, replacing energy which is lost as the body slows and/or falls during other parts of the stride. External work can be measured with fair approximation by means of a force plate, which records the horizontal and vertical components of the resultant force applied by the body to the ground over a complete stride. The horizontal force and the vertical force minus the body weight are integrated electronically to determine the instantaneous velocity in each plane. These velocities are squared and multiplied by one-half the mass to yield the instantaneous kinetic energy. The change in potential energy is calculated by integrating vertical velocity as a function of time to yield vertical displacement and multiplying this by body weight. The total mechanical energy as a function of time is obtained by adding the instantaneous kinetic and potential energies. The positive external mechanical work is obtained by adding the increments in total mechanical energy over an integral number of strides.

Kinematic Analysis of the Technique for Elite Male Long-Distance Speed Skaters in Curving

2007 ◽  
Vol 23 (2) ◽  
pp. 128-138 ◽  
Author(s):  
Jun Yuda ◽  
Masahiro Yuki ◽  
Toru Aoyanagi ◽  
Norihisa Fujii ◽  
Michiyoshi Ae
Keyword(s):  
Kinematic Analysis ◽  
High Speed ◽  
Center Of Mass ◽  
Three Dimensional ◽  
The Body ◽  
Long Distance ◽  
Video Cameras ◽  
World Class ◽  
Technical Factors ◽  
Support Leg

The purpose of this study was to investigate technical factors for maintaining skating velocity by kinematic analysis of the skating motion for elite long-distance skaters during the curve phase in official championship races. Sixteen world-class elite male skaters who participated in the 5,000-m race were videotaped with two synchronized high-speed video cameras (250 Hz) in a curve lane by using a panning DLT technique. Three-dimensional coordinates of the body and blades during the first and second halves of the races were collected to calculate kinematic parameters. In the group that maintained greater skating velocity, the thigh angle during the gliding phase of the left stroke during the second half was greater than that during the first half, and the center of mass was located more forward during the second half. Thus, it was suggested that long-distance speed skaters should change the support leg position during the gliding phase in the left stroke of the curve phase under fatigued conditions so that they could extend the support leg with a forward rotation of the thigh and less shank backward rotation.

Mechanical Energy Expenditures and Movement Efficiency in Full Body Reaching Movements

2010 ◽  
Vol 26 (1) ◽  
pp. 32-44 ◽  
Author(s):  
Daohang Sha ◽  
Christopher R. France ◽  
James S. Thomas
Keyword(s):  
Kinetic Energy ◽  
Potential Energy ◽  
Target Location ◽  
Mechanical Energy ◽  
The Body ◽  
Reaching Movements ◽  
Whole Body ◽  
Joint Power ◽  
Total Mechanical Energy ◽  
Full Body

The effect of target location, speed, and handedness on the average total mechanical energy and movement efficiency is studied in 15 healthy subjects (7 males and 8 females with age 22.9 ± 1.79 years old) performing full body reaching movements. The average total mechanical energy is measured as the time average of integration of joint power, potential energy, and kinetic energy respectively. Movement efficiency is calculated as the ratio of total kinetic energy to the total joint power and potential energy. Results show that speed and target location have significant effects on total mechanical energy and movement efficiency, but reaching hand only effects kinetic energy. From our findings we conclude that (1) efficiency in whole body reaching is dependent on whether the height of the body center of mass is raised or lowered during the task; (2) efficiency is increased as movement speed is increased, in part because of greater changes in potential energy; and (3) the CNS does not appear to use movement efficiency as a primary planning variable in full body reaching. It may be dependent on a combination of other factors or constraints.

Scanning electron microscopy of freeze-fractured prescapular lymph node of caprine

1984 ◽  
Vol 42 ◽  
pp. 244-245
Author(s):  
O. Faroon ◽  
F. Al-Bagdadi ◽  
T. G. Snider ◽  
C. Titkemeyer
Keyword(s):  
Lymph Node ◽  
Lymph Nodes ◽  
Lymphatic System ◽  
Three Dimensional ◽  
Meat Products ◽  
The Body ◽  
Morphological Data ◽  
The World ◽  
Cellular Constituent ◽  
Scanning Electron

The lymphatic system is very important in the immunological activities of the body. Clinicians confirm the diagnosis of infectious diseases by palpating the involved cutaneous lymph node for changes in size, heat, and consistency. Clinical pathologists diagnose systemic diseases through biopsies of superficial lymph nodes. In many parts of the world the goat is considered as an important source of milk and meat products.The lymphatic system has been studied extensively. These studies lack precise information on the natural morphology of the lymph nodes and their vascular and cellular constituent. This is due to using improper technique for such studies. A few studies used the SEM, conducted by cutting the lymph node with a blade. The morphological data collected by this method are artificial and do not reflect the normal three dimensional surface of the examined area of the lymph node. SEM has been used to study the lymph vessels and lymph nodes of different animals. No information on the cutaneous lymph nodes of the goat has ever been collected using the scanning electron microscope.

scholarly journals A biomechanical study of load carriage by two paired subjects in response to increased load mass

2021 ◽  
Vol 11 (1) ◽  
Author(s):  
Guillaume Fumery ◽  
Nicolas A. Turpin ◽  
Laetitia Claverie ◽  
Vincent Fourcassié ◽  
Pierre Moretto
Keyword(s):  
Recovery Rate ◽  
Energy Recovery ◽  
Mechanical Energy ◽  
Center Of Mass ◽  
Step Length ◽  
Biomechanical Study ◽  
Load Carriage ◽  
Joint Torque ◽  
Total Mechanical Energy ◽  
Load Mass

AbstractThe biomechanics of load carriage has been studied extensively with regards to single individuals, yet not so much with regards to collective transport. We investigated the biomechanics of walking in 10 paired individuals carrying a load that represented 20%, 30%, or 40% of the aggregated body-masses. We computed the energy recovery rate at the center of mass of the system consisting of the two individuals plus the carried load in order to test to what extent the pendulum-like behavior and the economy of the gait were affected. Joint torque was also computed to investigate the intra- and inter-subject strategies occurring in response to this. The ability of the subjects to move the whole system like a pendulum appeared rendered obvious through shortened step length and lowered vertical displacements at the center of mass of the system, while energy recovery rate and total mechanical energy remained constant. In parallel, an asymmetry of joint moment vertical amplitude and coupling among individuals in all pairs suggested the emergence of a leader/follower schema. Beyond the 30% threshold of increased load mass, the constraints at the joint level were balanced among individuals leading to a degraded pendulum-like behavior.

scholarly journals A faster escape does not enhance survival in zebrafish larvae

2017 ◽  
Vol 284 (1852) ◽  
pp. 20170359 ◽  
Author(s):  
Arjun Nair ◽  
Christy Nguyen ◽  
Matthew J. McHenry
Keyword(s):  
High Speed ◽  
Larval Fish ◽  
Body Position ◽  
Three Dimensional ◽  
Limited Range ◽  
The Body ◽  
Model Parameters ◽  
Fish Prey ◽  
Zebrafish Larvae ◽  
Fish Predators

An escape response is a rapid manoeuvre used by prey to evade predators. Performing this manoeuvre at greater speed, in a favourable direction, or from a longer distance have been hypothesized to enhance the survival of prey, but these ideas are difficult to test experimentally. We examined how prey survival depends on escape kinematics through a novel combination of experimentation and mathematical modelling. This approach focused on zebrafish ( Danio rerio ) larvae under predation by adults and juveniles of the same species. High-speed three-dimensional kinematics were used to track the body position of prey and predator and to determine the probability of behavioural actions by both fish. These measurements provided the basis for an agent-based probabilistic model that simulated the trajectories of the animals. Predictions of survivorship by this model were found by Monte Carlo simulations to agree with our observations and we examined how these predictions varied by changing individual model parameters. Contrary to expectation, we found that survival may not be improved by increasing the speed or altering the direction of the escape. Rather, zebrafish larvae operate with sufficiently high locomotor performance due to the relatively slow approach and limited range of suction feeding by fish predators. We did find that survival was enhanced when prey responded from a greater distance. This is an ability that depends on the capacity of the visual and lateral line systems to detect a looming threat. Therefore, performance in sensing, and not locomotion, is decisive for improving the survival of larval fish prey. These results offer a framework for understanding the evolution of predator–prey strategy that may inform prey survival in a broad diversity of animals.

Characteristics of Eight Irish Dance LandingsConsiderations for Training and Overuse Injury Prevention

2021 ◽  
Vol 25 (1) ◽  
pp. 30-37
Author(s):  
Sarah Klopp Christensen ◽  
Aaron Wayne Johnson ◽  
Natalie Van Wagoner ◽  
Taryn E. Corey ◽  
Matthew S. McClung ◽  
...  
Keyword(s):  
Large Range ◽  
Force Plate ◽  
Three Dimensional ◽  
The Body ◽  
Peak Force ◽  
Warm Up ◽  
Dance Training ◽  
Rise Rate ◽  
Dance Movements ◽  
Reaction Forces

Irish dance has evolved in aesthetics that lead to greater physical demands on dancers' bodies. Irish dancers must land from difficult moves without letting their knees bend or heels touch the ground, causing large forces to be absorbed by the body. The majority of injuries incurred by Irish dancers are due to overuse (79.6%). The purpose of this study was to determine loads on the body of female Irish dancers, including peak force, rise rate of force, and impulse, in eight common Irish hard shoe and soft shoe dance movements. It was hypothesized that these movements would produce different ground reac- tion force (GRF) characteristics. Sixteen female Irish dancers were recruited from the three highest competitive levels. Each performed a warm-up, reviewed the eight movements, and then performed each movement three times on a force plate, four in soft shoes and four in hard shoes. Ground reaction forces were measured using a three-dimensional force plate recording at 1,000 Hz. Peak force, rise rate, and vertical impulse were calculated. Peak forces normalized by each dancer's body weight for each of these variables were significantly different between move- ments and shoe types [F(15, 15)= 65.4, p < 0.01; F(15, 15) = 65.0, p < 0.01; and F(15, 15) = 67.4, p < 0.01, respectively]. The variable years of experience was not correlated with peak force, rise rate, or impulse (p > 0.40). It is concluded that there was a large range in GRF characteristics among the eight movements studied. Understanding the force of each dance step will allow instructors to develop training routines that help dancers adapt gradually to the high forces experienced in Irish dance training and competitions, thereby limiting the potential for overuse injuries.

scholarly journals Walking in simulated reduced gravity: mechanical energy fluctuations and exchange

1999 ◽  
Vol 86 (1) ◽  
pp. 383-390 ◽  
Author(s):  
Timothy M. Griffin ◽  
Neil A. Tolani ◽  
Rodger Kram
Keyword(s):  
Kinetic Energy ◽  
Potential Energy ◽  
Gravitational Potential ◽  
Inverted Pendulum ◽  
Mechanical Energy ◽  
Center Of Mass ◽  
Metabolic Cost ◽  
Metabolic Energy ◽  
Gravitational Potential Energy ◽  
Reduced Gravity

Walking humans conserve mechanical and, presumably, metabolic energy with an inverted pendulum-like exchange of gravitational potential energy and horizontal kinetic energy. Walking in simulated reduced gravity involves a relatively high metabolic cost, suggesting that the inverted-pendulum mechanism is disrupted because of a mismatch of potential and kinetic energy. We tested this hypothesis by measuring the fluctuations and exchange of mechanical energy of the center of mass at different combinations of velocity and simulated reduced gravity. Subjects walked with smaller fluctuations in horizontal velocity in lower gravity, such that the ratio of horizontal kinetic to gravitational potential energy fluctuations remained constant over a fourfold change in gravity. The amount of exchange, or percent recovery, at 1.00 m/s was not significantly different at 1.00, 0.75, and 0.50 G (average 64.4%), although it decreased to 48% at 0.25 G. As a result, the amount of work performed on the center of mass does not explain the relatively high metabolic cost of walking in simulated reduced gravity.

Many-legged maneuverability: dynamics of turning in hexapods

1999 ◽  
Vol 202 (12) ◽  
pp. 1603-1623 ◽  
Author(s):  
D.L. Jindrich ◽  
R.J. Full
Keyword(s):  
Horizontal Plane ◽  
Center Of Mass ◽  
Vertical Plane ◽  
Force Production ◽  
The Body ◽  
Mean Velocity ◽  
Blaberus Discoidalis ◽  
Body Design ◽  
Reaction Forces ◽  
Straight Ahead

Remarkable similarities in the vertical plane of forward motion exist among diverse legged runners. The effect of differences in posture may be reflected instead in maneuverability occurring in the horizontal plane. The maneuver we selected was turning during rapid running by the cockroach Blaberus discoidalis, a sprawled-postured arthropod. Executing a turn successfully involves at least two requirements. The animal's mean heading (the direction of the mean velocity vector of the center of mass) must be deflected, and the animal's body must rotate to keep the body axis aligned with the heading. We used two-dimensional kinematics to estimate net forces and rotational torques, and a photoelastic technique to estimate single-leg ground-reaction forces during turning. Stride frequencies and duty factors did not differ among legs during turning. The inside legs ended their steps closer to the body than during straight-ahead running, suggesting that they contributed to turning the body. However, the inside legs did not contribute forces or torques to turning the body, but actively pushed against the turn. Legs farther from the center of rotation on the outside of the turn contributed the majority of force and torque impulse which caused the body to turn. The dynamics of turning could not be predicted from kinematic measurements alone. To interpret the single-leg forces observed during turning, we have developed a general model that relates leg force production and leg position to turning performance. The model predicts that all legs could turn the body. Front legs can contribute most effectively to turning by producing forces nearly perpendicular to the heading, whereas middle and hind legs must produce additional force parallel to the heading. The force production necessary to turn required only minor alterations in the force hexapods generate during dynamically stable, straight-ahead locomotion. A consideration of maneuverability in the horizontal plane revealed that a sprawled-postured, hexapodal body design may provide exceptional performance with simplified control.

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