Memoirs: The Development of the Sabellid Branchiomma vesiculosum

1936 ◽  
Vol s2-78 (312) ◽  
pp. 543-603
Author(s):  
DOUGLAS P. WILSON

1. Larvae of Branchiomma vesiculosum Montagu were obtained from artificial fertilizations, and reared through metamorphosis and for some weeks afterwards. 2. The larvae are extremely yolky and do not feed until after metamorphosis. They swim by means of a broad prototroch, and are provided with a pair of cup-shaped eye-spots, a large head vesicle, and two anal vesicles. There is a broad neurotroch but no telotroch. The mouth is open but the anus is closed. In the last swimming stages rudiments of the adult branchial apparatus appear as a pair of lobed swellings, behind the eyes but in front of the prototroch. Behind the latter the collar rudiments appear. There are usually three or four chaetigerous segments marked out when metamorphosis sets in. 3. The metamorphosing larva settles on the bottom, and secretes for itself a tube of mucus. The prototroch and larval tissues (episphere) of the head clump together to form a large snout-like structure. This gradually breaks up into small pieces which are thrown off one by one as they are formed. With the loss of the prototroch and other larval head tissues the adult part of the head becomes joined on to the trunk. At the same time the branchial rudiments branch to form pinnules, which are directed forwards so that their bases overlap the eyes. The anus opens, and the young worm begins to feed. Metamorphosis occupies about four days. 4. The larval stages and the metamorphosis have been studied in histological, and to a limited extent in cytological, detail. Apart from the curious metamorphosis, which to some extent recalls that of Owenia and Polygordius, the development shows no striking features. 5. After metamorphosis the worm elongates by the addition of setigers in front of the pygidium. These are at first of the thoracic type (dorsal bristles, ventral uncini), but when the ninth and succeeding setigers appear they show abdominal constitution (dorsal uncini, ventral bristles) from the first. Thus no setigers change over from abdominal to thoracic constitution as they do in the development of young Serpulid worms. 6. Coincident with the loss of the neurotroch at metamorphosis the mid-dorsal line of the head and trunk of the young worm becomes ciliated to form the faecal groove of the adult. This strip of cilia continues on the ventral surfaces of the ninth and succeeding setigers, after passing round the right side between the eighth and ninth bristle segments. For a time the intersegmental groove between these segments is ciliated on the left side as well. 7. The branchial rudiments, which began to branch before metamorphosis, continue steadily to branch and grow afterwards. The pinnules are supported by an internal skeleton of thick-walled cells to the base of which the main dorsal and ventral longitudinal muscles of the body become attached. The most dorsal branch of each branchial rudiment, lying close to the mid-dorsal line, forms one of the so-called palps of the adult and is not supported by an internal skeleton. 8. The manner in which the young worms build their first sandy tube is described. 9. Early larvae of Sabella pavonina (Savigny) are described. They closely resemble those of Branchiomma. 10. The present position of embryological knowledge concerning the Polychaetes is very briefly summarized. It is shown that Sabellid larvae are more closely related to Serpulids than to those of any other family.

1891 ◽  
Vol s2-32 (128) ◽  
pp. 539-585
Author(s):  
FRANK E. BEDDARD

The principal new facts in this paper may be briefly summarised as follows : (1) The nephridial system consists of paired nephridia which do not open immediately on to the exterior, but are connected with an extensively ramifying system of tubes embedded in the circular and longitudinal muscular coats; these tubes consist of four principal longitudinal trunks continuous from segment to segment, and of a single large circular vessel in each segment passing right round the worm at the junction of the circular and longitudinal muscles; these are connected by a plexus of vessels, and numerous tubules, leading to the exterior, are given off from the circular trunk. In some of the genital segments the paired nephridia have almost disappeared, leaving only the integumental network. Nothing of the kind has been yet described in any Oligochæte. In the young worm, just escaped from the cocoon, there is no integumental network, which must, therefore, be regarded as secondary, but the anterior nephridia at any rate are connected on each side by a continuous longitudinal duct lying within the cœlom. (2) In the young worm the reproductive organs agree with these organs in other earthworms; in the^ adult, a large unpaired sac lying over the gut is developed; this sac encloses the receptacula ovorum, and opens by a median pore on Segment 13. It is developed from mesoblastic tissues, and is not therefore the morphological equivalent of the spermathecæ in Lumbricus, &c., but it performs the same function; the sac is formed internally and then grows out towards the epidermis ; it is at first in open communication with the cœlom; its front wall is formed out of the intersegmental septum between Segments 12, 13; the ovaries are enclosed by it, but disappear early, before the sac is completed; otherwise the ova would be probably unable to enter the egg-sac which becomes nearly completely shut off from the sac; the two are in communication only by the oviducal funnel, which has become divided by the growth of the spermathecal sac into two separate tubes, one opening into the spermathecal sac, the other into the closed egg-sac; they unite, of course, to form the oviduct itself, which opens on to the 15th segment, reckoning by the external furrow, but on to the border line between Segments 14, 15, reckoning by the septa. (3) The testes and the vas deferens funnels are quite typical in their structure and position; so, too, are the (two) pairs of sperm-sacs (in Segments 11, 12). The sperm ducts are not, as they are in other Eudrilidæ, dilated to form sperm reservoirs ; they open into tubular atria, with thick muscular walls and glandular lining, near to their blind extremities; the two atria open by a common pore upon the border line between Segments 17, 18; each is furnished with a short penial seta not ornamented. (4) The alimentary tract has no calciferous glands or ventral œsophageal pouches such as are found in other Eudrilidæ at the end of the oesophagus are three gizzards, one to a segment; the intestine which immediately follows has at first three typhlosolar folds; later on the two lateral and shorter folds disappear. The ventral wall of the pharynx is connected with the nephridial tubes of its segments; they open into the interior of the pharynx. (5) The area surrounding the setas of each side of the body is shut off from the general body-cavity, forming a paired series of chambers; in the œsophageal region is developed a perihæmal cœlomic space surrounding the subœsophageal vessels.


1902 ◽  
Vol s2-46 (181) ◽  
pp. 97-141
Author(s):  
ISABELLA M. DRUMMOND

To sum up, then, theories of Gasteropod torsion may be divided into two classes: a. Those which view the present position of the palleal complex as due to a forward movement along the right side of the body, which resulted from greater growth of the left side of the body than of the right. h. Those which view the present position of the palleal complex as due to a ventral flexion followed by a vertical rotation of the whole visceral hump upon the head. The evidence for the second of these views seems greater than that for the first, in that-- 1. A vertical displacement through 180° of all the organs contained in the visceral hump takes place in the course of ontogeny. 2. There is some evidence, both from comparative anatomy and embryology, for believing that the oesophagus has undergone an actual twist. 3. Monstrosities which retain the palleal complex in a ventral position show a tendency to form an exogastric coil. The innervation of the mantle was shown to be equally difficult to explain on either hypothesis. Also, against the first view was urged the insufficiency of the evidence upon which Bütschli bases his conclusions with regard to zones of unequal growth. With regard to the phylogenetic cause of the vertical twist, embryology can only give negative evidence; while in considering the ontogenetic cause we are thrown back upon unsolved problems of heredity, and must confess our ignorance. In conclusion, I wish to offer most hearty thanks to Professor Weldon, not only for having placed freely at my disposal all the resources of the laboratory, but also for most kind personal aid at all stages of the work. My thanks are also due to Mr. Richard Evans for much help in the technique.


1984 ◽  
Vol 35 (3) ◽  
pp. 315 ◽  
Author(s):  
RA Rose ◽  
TG Dix

Planktotrophic larvae of C. asperrimus were reared from zygotes with a mean diameter of 61.5�m. The smallest straight-hinged larval shell was 80�m long, and slightly protruding umbones were formed 6 days after fertilization. The length (L) and height (H) relationship of larval shells was described by the allometric growth curve H = 0.309 L1 203. Provincular structure of 13-day-old larval shells was simple and there were nine taxodontal teeth per valve, four at one end of the hinge line and five at the other end. Eye spots were conspicuous and common in pediveligers 160-220�m long. Metamorphosis occurred 20-22 days after fertilization at 17-18�C when larvae were 190�m or longer. Larvae settled on monofilament nylon collectors. Newly settled juveniles developed a distinctive byssal notch on the right valve but attachment by byssal threads was never permanent. The small size of eggs and early larvae and allometric growth of prodissoconch II shells are the most significant characteristics distinguishing the larval stages of C. asperrimus from other pectinids in Tasmanian waters.


Author(s):  
Anne Phillips

No one wants to be treated like an object, regarded as an item of property, or put up for sale. Yet many people frame personal autonomy in terms of self-ownership, representing themselves as property owners with the right to do as they wish with their bodies. Others do not use the language of property, but are similarly insistent on the rights of free individuals to decide for themselves whether to engage in commercial transactions for sex, reproduction, or organ sales. Drawing on analyses of rape, surrogacy, and markets in human organs, this book challenges notions of freedom based on ownership of our bodies and argues against the normalization of markets in bodily services and parts. The book explores the risks associated with metaphors of property and the reasons why the commodification of the body remains problematic. The book asks what is wrong with thinking of oneself as the owner of one's body? What is wrong with making our bodies available for rent or sale? What, if anything, is the difference between markets in sex, reproduction, or human body parts, and the other markets we commonly applaud? The book contends that body markets occupy the outer edges of a continuum that is, in some way, a feature of all labor markets. But it also emphasizes that we all have bodies, and considers the implications of this otherwise banal fact for equality. Bodies remind us of shared vulnerability, alerting us to the common experience of living as embodied beings in the same world. Examining the complex issue of body exceptionalism, the book demonstrates that treating the body as property makes human equality harder to comprehend.


2013 ◽  
Vol 7 (1) ◽  
pp. 7
Author(s):  
Doni Budiono

The  authority  of justice in Indonesia  is executed by  the Supreme Courts and  the  justice  boards/body under the Supreme Courts, including  the general  justice, religious affairs justice, military justice,  state administration  justice,  and  the Constitution Court. According to  certainty in  the Act of  Tax Court, Article1, clause  (5),  tax  dispute   refers to the legal dispute arising in the  taxation  affairs between the  tax payer or the  body  responsible for the  tax with   the government   executives  ( Directorate General of Tax) as the consequence of   the issue of  the decree for the  appeal  to the Tax  Court in accordance with the  tax Act, including the  charge  against the  execution of collection   in accordance with the  Act of Tax Collection by force. The  formation of Tax Court is  designed by  the Executives, in this case, the  Department of Finance, specifically  the Directorate   General  of Tax  which has the right to issue  law  more technical about  tax accord to Article 14,  letter A,  President Decree  no. 44  year 1974,  concerning the  basic  organization of the Department.  Based on  it,  it  is clear that  in addition to execute the government  rules and policy,  this body  has to execute judicial   rules and policy. This is against the  principles of  Judicative  Power/Authority in Indonesia,  which   clearly states that this body  should be under the Supreme Court.   Therefore. It is suggested that   the Act  No UU no.14 Year 2012 concerning  Tax Court   be revised  in accordance with the system of  Power Division  of Justice  as  stated in 45 Constitutions.


2020 ◽  
Vol 11 (SPL1) ◽  
pp. 1477-1481
Author(s):  
Ishwari Gaikwad ◽  
Priyanka Shelotkar

The current world situation is both frightening and alarming due to the massive disruption caused by the Covid-19 pandemic. The next few days are censorious as we need to be very precautious in our daily regimen as well as dietary habits. Ayurveda offers knowledge about food based on certain reasoning. Indecent food custom is the chief cause for the rising development of health disorders in the current era. In classical texts of Ayurveda, the concept of diet explained well, ranging from their natural sources, properties and specific utility in pathological as well as physiological manner. In this work, the review of the relevant literature of Ahara (Diet) was carried out from Charak Samhita and other texts, newspapers, articles, web page related to the same.  Every human being is unique with respect to his Prakriti (Physical and mental temperament), Agni (Digestive capacity), Koshtha  (Nature of bowel) etc. For that reason, the specificity of the individual should be kept in mind. Ahara, when consumed in the appropriate amount at the right moment following all Niyamas (Guidelines) given in Ayurveda texts, gives immunity and keeps the body in a healthy state during pandemics such as Covid-19. Ultimately, this will help the human body to maintain its strength for life. This article reviews the concept of diet viz. combination of foods, their quantity and quality, methods of preparation and processing, which are to be followed during pandemics and are essential in maintenance and endorsement of health and preclusion of diseases.


2020 ◽  
Vol 11 (SPL1) ◽  
pp. 832-838
Author(s):  
Roshna Sukheoji Bhutada ◽  
Renu Rathi ◽  
Devyani Dasar

WHO declared Covid 19 /SARS -COV-2 as a global pandemic.Till date, there is no medicine for COVID-19. If the Infection arises in the body then the defence mechanism activated against infection. A recent study suggests that temporarily augmenting the body's immune system in the early stages of COVID-19 can help patient to avoid severe symptoms as it is rightly said prevention is better than cure. Ayurveda approaches to develop physiological reactions to facilitate immunity. Planning of diet is most important to boost immunity.As per many researches to provide supplementary food which contains Zinc, Vitamin C,Vitamin D and immunity boosting foodsuch as citrus natural products, custard apple, apple, papaya is among the Fruits. Vegetables include broccoli, onion, garlic and green leafy vegetables. Nuts, ginger, turmeric, pepper, egg yolk, shellfish, mushroom. The need of the hour is a quick boost to immune system to keep it fit, fighting. One should get the right amount of nutrients from the diet, supplementation regimen to boost immune system.In this review, there are few common supplements and super food studies have been included. It might be a torch bearer as sample menu and their alternatives are given for a normal adult. Needy may change contemplated according to age, sex, body mass index and daily physical activities.


2020 ◽  
Vol 324 (2) ◽  
pp. 242-251
Author(s):  
L.P. Flyachinskaya ◽  
P.A. Lezin

The paper considers the development of Ciliatocardium ciliatum from the stage of straight hinge to juvenile. In the White Sea the spawning of C. ciliatum begins at the end of June, larvae at different stages of development occur in plankton until the end of September. The earliest of the larvae found had shell lengths of 123–130 µm. The paper first examined the anatomy and structure of the larval shell of C. ciliatum. During the development, the main stages of organogenesis were described and special attention was paid to the formation of the digestive and muscular systems. The digestive system begins to function when the larva reaches a size of 170–180 µm. The digestive gland has a two-blade shape and is shifted to the right side. The foot is formed at a size of 230 µm, the gill rudiments appear when the larva reaches 270 µm. The development of the larval shell and larval hinge of the mollusc is considered in detail. The development of the larval shell of C. ciliatum is similar to the development of other family members. Throughout all the larval stages, the shell has a rounded shape with a low umbos, and the prodissoconch II has a clearly visible concentric structure. The C. ciliatum larval hinge is characterized by weak differentiation and the absence of pronounced cardinal teeth typical for other Cardiidae. However, the lateral structures of the castle – ridges and flanges – are well developed. The ligament begins to form at a size of 240–250 µm and occupies a lateral position. The settlement of the cockle takes place in September in the subtidal zone. After the metamorphosis, a large radial sculpture is formed on the dissoconch and a number of small spikes are formed at the rib of the posterior shoulder.


Impact ◽  
2018 ◽  
Vol 2018 (3) ◽  
pp. 26-28
Author(s):  
Jonathan Dawson ◽  
Richard Oreffo

Gels made from clay could provide an environment able to stimulate stem-cells due to their ability to bind biological molecules. That molecules stick to clay has been known by scientists since the 1960s. Doctors observed that absorption into the blood stream of certain drugs was severely reduced when patients were also receiving clay-based antacid or anti-diarrhoeal treatments. This curious phenomenon was realized to be due to binding of the drugs by clay particles. This interaction is now routinely harnessed in the design of tablets to carefully control the release and action of a drug. Dr Dawson now proposes to use this property of clay to create micro-environments that could stimulate stem cells to regenerate damaged tissues such as bone, cartilage or skin. The rich electrostatic properties of nano (1 millionth of a millimetre) -scale clay particles which mediate these interactions could allow two hurdles facing the development of stem-cell based regenerative therapies to be overcome simultaneously. The first challenge - to deliver and hold stem cells at the right location in the body - is met by the ability of clays to self-organise into gels via the electrostatic interactions of the particles with each other. Cells mixed with a low concentration (less than 4%) of clay particles can be injected into the body and held in the right place by the gel, eliminating, in many situations, the need for surgery. Clay particles can also interact with large structural molecules (polymers) which are frequently used in the development of materials (or 'scaffolds'), designed to host stem cells. These interactions can greatly improve the strength of such structures and could be applied to preserve their stability at the site of injury until regeneration is complete. While several gels and scaffold materials have been designed to deliver and hold stem cells at the site of regeneration, the ability of clay nanoparticles to overcome a second critical hurdle facing stem-cell therapy is what makes them especially exciting. Essential to directing the activity of stem-cells is the carefully controlled provision of key biological signalling molecules. However, the open structures of conventional scaffolds or gels, while essential for the diffusion of nutrients to the cells, means their ability to hold the signalling molecules in the same location as the cells is limited. The ability of clay nano-particles to bind biological molecules presents a unique opportunity to create local environments at a site of injury or disease that can stimulate and control stem-cell driven repair. Dr Dawson's early studies investigated the ability of clay gels to stimulate the growth of new blood vessels by incorporating a key molecular signal that stimulates this process, vascular endothelial growth factor (VEGF). In a manner reminiscent of the observations made in the 60s, Dr Dawson and colleagues observed that adding a drop of clay gel to a solution containing VEGF caused, after a few hours, the disappearance of VEGF from the solution as it became bound to the gel. When placed in an experimental injury model, the gel-bound VEGF stimulated a cluster of new blood vessels to form. These exciting results indicate the potential of clay nanoparticles to create tailor-made micro-environments to foster stem cell regeneration. Dr Dawson is developing this approach as a means of first exploring the biological signals necessary to successfully control stem cell behaviour for regeneration and then, using the same approach, to provide stem cells with these signals to stimulate regeneration in the body. The project will seek to test this approach to regenerate bone lost to cancer or hip replacement failure. If successful the same technology may be applied to harness stem cells for the treatment of a whole host of different scenarios, from burn victims to those suffering with diabetes or Parkinson's.


2021 ◽  
pp. 1-20
Author(s):  
Benjamin Hegarty

The regulation of public space is generative of new approaches to gender nonconformity. In 1968 in Jakarta, the capital of Indonesia, a group of people who identified as wadam—a new term made by combining parts of Indonesian words denoting “femininity” and “masculinity”—made a claim to the city's governor that they had the right to appear in public space. This article illustrates the paradoxical achievement of obtaining recognition on terms constituted through public nuisance regulations governing access to and movement through space. The origins and diffuse effects of recognition achieved by those who identified as wadam and, a decade later, waria facilitated the partial recognition of a status that was legal but nonconforming. This possibility emerged out of city-level innovations and historical conceptualizations of the body in Indonesia. Attending to the way that gender nonconformity was folded into existing methods of codifying space at the scale of the city reflects a broader anxiety over who can enter public space and on what basis. Considering a concern for struggles to contend with nonconformity on spatial grounds at the level of the city encourages an alternative perspective on the emergence of gender and sexual morality as a definitive feature of national belonging in Indonesia and elsewhere.


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