scholarly journals Wolf Howling and Its Role in Territory Maintenance

Behaviour ◽  
1979 ◽  
Vol 68 (3-4) ◽  
pp. 207-249 ◽  
Author(s):  
Fred H. Harrington ◽  
L. David Mech

AbstractAn experimental study of the role of howling in wolf territory maintenance was conducted in the Superior National Forest, Minnesota. Vocal replies and behaviour of radio-collared wolves in response to human howls were analyzed for eight packs and 10 lone wolves during a 2-year period. Reply rate varied significantly throughout the year. A mid-winter increase was correlated with the breeding season, especially for groups containing breeding animals (alpha male or alpha female). A second, longer increase in reply rate started in midsummer, peaked about August, and declined to a low in early winter. The decline in autumn howling response occurred sooner in a pack whose pups developed faster. Through the year, the howling reply rate was significantly higher among all packs and lone wolves attending prey kills. The more food remaining at a kill, the higher the reply rate was. For wolves separated from their pack, the howling reply rate was dependent on their age and social role. Among adults, only alpha males ever replied alone, and their reply rate, and number of howls per session, exceeded those of other animals. Alpha males sometimes approached during howling sessions, whereas other adults usually retreated. Younger animals replied more often as pups than as yearlings, and then only during their first 7 months, after which they replied little more than most adults. Finally, larger packs replied more often than smaller packs. Specific behaviours noted during howling sessions, including movements away from the howler, indicated that howling was related to interpack agonism. In addition, three of the major factors influencing reply rate also significantly affect the level of agonism toward pack strangers : pack size, social role, and breeding season. The other two factors, kills and pups, are both important pack resources necessitating exclusive occupancy of a site. The high reply rates at sites containing kills or pups constitute strong circumstantial evidence that howling is important in territory maintenance. During howling sessions, wolves usually remained near their original site after replying, or retreated if they remained silent. This difference apparently was related to the problem of avoiding both accidental and deliberate encounters, and to cost/benefit considerations at the wolves' location. Howling was considered most effective in mediating avoidance in two situations : when two packs approached a common area of overlap, and when a pack returned to an area little used for weeks, in which scent posts would have lost effectiveness in deterring strangers. Both scent-marking and howling apparently are important in spacing. However, they differ in their roles and are complementary, with scent-marking being long-term and site-specific, and howling being immediate and long-range. Finally, lone wolves which do not possess territories, rarely replied, sharing the "low-profile" behaviour expected of surplus animals in a territorial population.

2018 ◽  
Vol 103 (6) ◽  
pp. 781-788 ◽  
Author(s):  
Geetha Iyer ◽  
Bhaskar Srinivasan ◽  
Shweta Agarwal ◽  
Ruchika Pattanaik ◽  
Ekta Rishi ◽  
...  

PurposeTo analyse the functional and anatomical outcomes of different types of keratoprostheses in eyes with retained silicone oil following vitreoretinal surgery.MethodsRetrospective chart review of patients operated with any type of permanent keratoprosthesis (Kpro) in silicone oil-filled eyes between March 2003 and June 2017 were analysed.Results40 silicone oil-filled eyes underwent keratoprostheses, of which 22 were type 1 and 18 were type 2 Kpros (Lucia variant—nine, modified osteo odonto kerato prosthesis (MOOKP)—four, Boston type 2—three and osteoKpro—two) with a mean follow-up of 61.54 , 42.77, 45.25 , 25 and 37 months, respectively. Anatomic retention of the primary Kpro was noted in 33 eyes (82.5%). A best-corrected visual acuity of better than 20/200 and 20/400 was achieved in 26 (65%)+32 (80%) eyes. Retroprosthetic membrane (RPM) was the most common complication noted in 17 eyes (42.5%). Perioptic graft melt was noted in 4 of 22 eyes of the type 1 Kpro (2 (10.5%) without associated ocular surface disorder (OSD)) and in 1 eye each of Boston and Lucia type 2 Kpro. Laminar resorption occurred in one eye each of the MOOKP and OKP groups. Endophthalmitis and glaucoma did not occur in any eye.ConclusionAppropriately chosen keratoprosthesis is a viable option for visual rehabilitation in eyes post vitreoretinal surgery with retained silicone oil-induced keratopathy not amenable to conventional penetrating keratoplasty. Kpro melt among type 1 Kpro did not occur in 89.5% eyes without associated OSD (19 of 22 eyes), despite the lack of aqueous humour and presence of RPM (4 eyes), two factors considered to play a significant role in the causation of sterile melts. Of interest to note was the absence of infection in any of these eyes. The possible protective role of oil from endophthalmitis is interesting, though yet to be ascertained.


2018 ◽  
Vol 1 (2) ◽  
pp. 101
Author(s):  
Vesna Srnic ◽  
Emina Berbic Kolar ◽  
Igor Ilic

<p><em>In addition to the well-known classification of long-term and short-term memory, we are also interested in distinguishing episodic, semantic and procedural memory in the areas of linguistic narrative and multimedial semantic deconstruction in postmodernism. We compare the liveliness of memorization in literary tradition and literature art with postmodernist divisions and reverberations of traditional memorizations through human multitasking and performative multimedia art, as well as formulate the existence of creative, intuitive and superhuman paradigms.</em></p><em>Since the memory can be physical, psychological or spiritual, according to neurobiologist Dr. J. Bauer (Das Gedächtnis des Körpers, 2004), the greatest importance for memorizing has the social role of collaboration, and consequently the personal transformation and remodelling of genomic architecture, yet the media theorist Mark Hansen thinks technology brings different solutions of framing function (Hansen, 2000). We believe that postmodern deconstruction does not necessarily damage memory, especially in the field of human multitasking that utilizes multimedia performative art by means of anthropologization of technology, thereby enhancing artistic and affective pre&amp;post-linguistic experience while unifying technology and humans through intuitive empathy in society.</em>


Author(s):  
Mireille Rosello

This particular attempt at imagining a site of memory made of words may appear irreverent at first, but it has been crafted as an homage to a formidable woman: Jeanne Duval. I have taken the liberty of fictionalizing a first-person narrator who will talk about ‘herself’, at the risk of usurping her voice and her identity. Jeanne (whose name was or was not Duval) was a woman of colour and she had a long-term turbulent relationship with the enfant terrible of French nineteenth-century poetry, Charles Baudelaire. As a result, historical accounts both magnify and marginalize her. Trying to do justice to a historical character who was so much more than a muse but may not have been happy to embrace the role of exemplary black foremother, this text puts together the numerous and often incompatible portraits of Jeanne Duval. She appears and disappears in biographies (Emmanuel Richon), novels (Fabienne Pasquet), short stories (Angela Carter), academic studies (Claude Pichois). She is both present and absent, celebrated and erased in the so-called ‘Black Venus cycle’ of Baudelaire’s Flower of Evil as well as in paintings by Edouard Manet (Baudelaire’s Mistress, Reclining) and Gustave Courbet (The Painter’s Studio). The objective was to question the process of memorialization that might silence or appropriate her instead of providing her with a safe space of memory. It remains to be seen to what extent Jeanne is here celebrated or betrayed.


2006 ◽  
Vol 36 (1) ◽  
pp. 23-49 ◽  
Author(s):  
Goolam Vahed

AbstractThis study focuses on Durban's Grey Street mosque, built by Indian Memon migrants in 1880. This review of the first half-century of the mosque's existence underlines the important social role of mosques, and also questions the notion of homogeneous Muslim community. While the mosque was the most visible symbol of Muslim identity in Natal, it was also a site of contestation, reflecting the class, language, caste and ethnic divisions among Muslims in a diasporic situation. Mosques were built along class and ethnic lines and dominated by traders. As Muslim society matured, there were challenges to the leadership of non-clerical traders who did not tolerate challenges to their authority. Opposition sometimes centred on Imams who commanded the allegiance of the congregation. Mosques did not have an independent life but reflected the prevailing power structures in Muslim society. While outsiders believed that ethnic diversity was subsumed by a unitary Muslim mass, Muslims comprised a community of communities, and the building and management of mosques underlined this fact.


Author(s):  
K. S. Tarasov

This article analyses discussions between representatives of three schools in the theory of international relations - realism, liberalism and constructivism - on the driving factors of nuclear proliferation. The paper examines major theoretical approaches, outlined in the studies of Russian and foreign scientists, to the causes of nuclear weapons development, while unveiling their advantages and limitations. Much of the article has been devoted to alternative approaches, particularly, the role of mathematical modeling in assessing proliferation risks. The analysis also reveals a variety of different approaches to nuclear weapons acquisition, as well as the absence of a comprehensive proliferation theory. Based on the research results the study uncovers major factors both favoring and impeding nuclear proliferation. The author shows that the lack of consensus between realists, liberals and constructivists on the nature of proliferation led a number of scientists to an attempt to explain nuclear rationale by drawing from the insights of more than one school in the theory of IR. Detailed study of the proliferation puzzle contributes to a greater understating of contemporary international realities, helps to identify mechanisms that are most likely to deter states from obtaining nuclear weapons and is of the outmost importance in predicting short- and long-term security environment. Furthermore, analysis of the existing scientific literature on nuclear proliferation helps to determine future research agenda of the subject at hand.


2020 ◽  
Author(s):  
Aditya Ghoshal ◽  
Anuradha Bhat

AbstractMate search tactics and association preferences among organisms in natural habitats can be dynamic and are determined by inherent trait preferences as well as the cost-benefit trade-offs associated with each mating decision. Two of the prime factors regulating mating decisions are the presence of competing conspecifics and predatory threats, both of which have important fitness consequences for the individual. We studied the influence of these two factors separately in mate search tactics and association preferences among zebrafish males. Male zebrafish were presented with a choice of two patches, consisting of different number of females, of which one patch was also associated with a predatory threat. We found that males made a preferential choice for the patch with more number of females only when the numerical difference between choices are starkly different, irrespective of the predatory threat associated with the patch. This points towards the role of numerical cognition in assessing cost-benefit tradeoffs in male zebrafish. We also studied the association preference of males in a multi-choice setup, consisting of four separate mixed-sex groups of zebrafish varying in densities. Our results showed that while test males preferred to visit the male-biased patches more often, they spent more time near female-biased patches or patches with equal sex ratio patches indicating the role of complex interplay of social cues in determining the associative behavior of males to a patch. This study, thus, sheds further light on the interactive roles of social cues and cognitive abilities in mate association patterns in this species.


Author(s):  
Pierre Pestieau ◽  
Mathieu Lefebvre

This chapter is concerned with the rise in long-term care needs. Long-term care concerns individuals who are no longer able to carry out basic daily activities. Most of the care is currently provided by informal caregivers, mainly the family, while the role of formal care provided by the state or the market remains small. The chapter explains, however, why informal care is expected to decline and analyses the low private insurance development, the so-called long-term care insurance puzzle. These two factors, the decreasing role of the family and a thin insurance market, plead for the development of a full fledge social insurance for long-term care. The chapter then looks at the optimal design of such an insurance.


Oryx ◽  
2018 ◽  
Vol 52 (4) ◽  
pp. 709-717 ◽  
Author(s):  
Luca Canova ◽  
Alessandro Balestrieri

AbstractWe monitored egg clutch numbers of a population of the endemic Italian agile frog Rana latastei in a Site of Community Interest in northern Italy (SCI IT 209000) during 1997–2017 with the aim of assessing the long-term variation in its abundance. We walked along the banks of canals and small ponds (n = 22) 1–3 times per week between early February and mid-April each year to detect egg clutches. The relationships between the start of the breeding season, yearly egg mass counts, rate of yearly change in the number of recorded egg masses and 15 climatic and environmental variables were assessed by multiple regression. The first deposition of eggs occurred progressively later in the year throughout the study period and mean air temperature during the breeding season decreased over this period. Agile frogs showed high deposition site-fidelity. Despite large variations in the number of egg clutches detected from year to year, the population size remained stable in the long term. Peaks in the number of egg clutches occurred 2 years after the dredging of canals, carried out every 4–6 years to improve water availability, starting in 2004 as part of a LIFE Nature Project. This was the only predictor of the number of egg clutches deposited, suggesting that periodical management is needed to support the agile frog population. Our results reinforce the need for multi-year monitoring to determine both the long-term success of habitat restoration projects and the status of residual populations.


2008 ◽  
Vol 9 (1) ◽  
pp. 1-16 ◽  
Author(s):  
Chuan-Zhong Li ◽  
Karl-Gustaf Löfgren

Abstract This paper deals with the modern theory of social cost-benefit analysis in a dynamic economy. The theory emphasizes the role of a comprehensive, forward looking, dynamic welfare index within the period of the project rather than that of a project’s long-term consequences. However, what constitutes such a welfare index remains controversial in the recent literature. In this paper, we attempt to shed light on the issue by deriving three equivalent cost-benefit rules for evaluating a small project. In particular, we show that the direct change in a net national product (NNP) qualifies as a convenient welfare index without involving any other induced side effects. The project evaluation criterion thus becomes the present discounted value of the direct changes in NNP over the project period. We also illustrate the application of this theory in a few stylized examples.


PLoS ONE ◽  
2021 ◽  
Vol 16 (12) ◽  
pp. e0260728
Author(s):  
Carlota Pagès-Portabella ◽  
Mila Bertolo ◽  
Juan M. Toro

In western music, harmonic expectations can be fulfilled or broken by unexpected chords. Musical irregularities in the absence of auditory deviance elicit well-studied neural responses (e.g. ERAN, P3, N5). These responses are sensitive to schematic expectations (induced by syntactic rules of chord succession) and veridical expectations about predictability (induced by experimental regularities). However, the cognitive and sensory contributions to these responses and their plasticity as a result of musical training remains under debate. In the present study, we explored whether the neural processing of pure acoustic violations is affected by schematic and veridical expectations. Moreover, we investigated whether these two factors interact with long-term musical training. In Experiment 1, we registered the ERPs elicited by dissonant clusters placed either at the middle or the ending position of chord cadences. In Experiment 2, we presented to the listeners with a high proportion of cadences ending in a dissonant chord. In both experiments, we compared the ERPs of musicians and non-musicians. Dissonant clusters elicited distinctive neural responses (an early negativity, the P3 and the N5). While the EN was not affected by syntactic rules, the P3a and P3b were larger for dissonant closures than for middle dissonant chords. Interestingly, these components were larger in musicians than in non-musicians, while the N5 was the opposite. Finally, the predictability of dissonant closures in our experiment did not modulate any of the ERPs. Our study suggests that, at early time windows, dissonance is processed based on acoustic deviance independently of syntactic rules. However, at longer latencies, listeners may be able to engage integration mechanisms and further processes of attentional and structural analysis dependent on musical hierarchies, which are enhanced in musicians.


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